Belzebub, Vereshchaka & Olesen & Lunina, 2016

Vereshchaka, Alexander L., Olesen, Jørgen & Lunina, Anastasia A., 2016, A phylogeny-based revision of the family Luciferidae (Crustacea: Decapoda), Zoological Journal of the Linnean Society 178 (1), pp. 15-32 : 26-28

KEY TO SPECIES OF BELZEBUB GEN. NOV.

1. Petasma: sheath bottle-shaped, with distal transverse rows of ridges, plate-like structures large and fringed ( Fig. 5C, D ) ……. B. intermedius ( Hansen, 1919) comb. nov.

– Petasma: sheath subconical, with distal longitudinal rows of fine scales, plate-like structures small, with entire margins ( Fig. 7F ) ……… 2

2. Petasma: processus ventralis bearing a dense brush of long terminal setae ( Fig. 6D – F ) ………. B. penicillifer ( Hansen, 1919) comb. nov.

– Petasma: processus ventralis with short, scattered lateral setae only, without dense brush of terminal setae ( Figs 7F , 8D ) …….. 3

3. Outer tooth of uropodal exopod far from reaching distal end of exopod; petasma: processus ventralis setose along medial part ( Fig. 7G, F ) …… B. hanseni ( Nobili, 1905) comb. nov.

– Outer tooth of uropodal exopod nearly reaching distal end of exopod; petasma: processus ventralis setose along distal part ( Fig. 8D ) …….. 4

4. Atlantic species …… B. faxoni ( Borradaile, 1915) comb. nov.

– Indo-Pacific species ………. B. chacei ( Bowman & McCain, 1967) comb. nov.

Figures

Figure 5. Petasma of Belzebub intermedius (Hansen, 1919) comb. nov.: general structure (A), apical view of the sheath (B, scanning electron micrograph), lateral view of the sheath and the processus ventralis (C, D, scanning electron micrographs). Arrows point at same morphological structures.

Figure 6. Petasma of Belzebub peпicillifer (Hansen, 1919) comb. nov.: general structure (A), apical view of the petasma (B, scanning electron micrograph), apical view of the sheath (C, scanning electron micrograph), lateral view of the sheath and the processus ventralis (D, E, scanning electron micrographs), apex of the processus ventralis (F, scanning electron micrograph). Arrows point at same morphological structures.

Figure 7. Petasma of Belzebub hanseni (Nobili, 1905) comb. nov.: ‘thumb’ (A, scanning electron micrograph), general structure (B), apical view of the petasma (C, scanning micrograph), lateral view of the sheath and the processus ventralis (D, scanning electron micrograph, E), apex of processus ventralis and the sheath (F, G, scanning electron micrographs). Arrows point at same morphological structures.

Figure 8. Petasma of Belzebub faxoni (Borradaile, 1915) comb. nov.: ‘thumb’ (A, scanning electron micrograph), general structure (B), apical view of the petasma (C, scanning electron micrograph), lateral view of the sheath and the processus ventralis (D, scanning electron micrograph, E, scanning electron micrograph, F), distal part of the sheath with apical scales (G–I, scanning electron micrographs). Arrows point at same morphological structures.