Cochlostoma (T.) arnautorum ( Wagner, 1906 )

Zallot, Enrico, Kamchev, Panche, Schilthuizen, Menno, Fehér, Zoltán, Mattia, Willy De & Gittenberger, Edmund, 2024, Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae), European Journal of Taxonomy 927, pp. 1-163 : 129-131

Cochlostoma (T.) arnautorum ( Wagner, 1906)

Figs 105 (yellow dots), 112–113

Auritus (Auritus) kleciaki arnautorum Wagner, 1906: 131–132 , pl. 4 fig. 17a–b.

Lectotype (here designated)

ALBANIA • ♀; “ Auritus Kleciaki arnautorum A. J. Wgn., Wagner det., Selci , Montenegro , ex Coll. Wohlberedt”; NHMW35442 .

Other specimens

ALBANIA • 1- Selcë (topotypical); 42.5199° N, 19.6271° E; 2018; M. Schilthuizen and I. Njunjić leg.; TxExDU 0137 .

Type locality

The author ( Wagner 1906: 131–132) listed a series of localities: “Rikovak, Selci (“Selci” at that time in Montenegro is now Selcë, Albania) und Korito in Montenegro; Dr. Sturany sammelte entsprechende Exemplare zwischen Fundina und Zatriebac an der montenegrinisch-albanesischen Grenze, ferner eine kleinere, dunkelrotbraun gefärbte Form mit etwas kräftigeren und weitläufigeren Rippchen auf den oberen Umgängen am Berge Sildinja östlich von Skutari in Albanien.”. Due to the present lectotype designation, the type locality is restricted to ALBANIA • Shkodër County, Selcë .

Description

SHELL. Widely spaced riblets on last part of protonch. Teleoconch spotless, ribbed with rather prominent ribs. Moderately strong lip with abrubtly inwardly curved columellar lobe covering umbilicus.

MEASUREMENTS. 6 ♀♀: whorls= 7.4–8.5, H = 6–8 mm, H/W= 2.46–2.81, roundness =0.12–0.17, ribs incl.= 57–68°, apert. incl.= 16–29°, ribs/mm 1 st wh.= 7–12, ribs/mm 4 th wh.=7–11.

FEMALE GENITAL ORGANS. Posterior connection of pedunculus to bursa copulatrix. Seminal receptacle with indistinct basis and apex, almost tubular, with 2–3 narrow loops close to apex. Junction of uterus gland close to distal oviduct-pedunculus connection.

Remarks

Wagner (1906) described this taxon as a subspecies of C. (T.) kleciaki , and as such it is reported in MolluscaBase (2023). We consider it a separate species because of the differences in the female genital morphology: the junction of the uterus gland is close to the connection between the distal oviduct and the pedunculus of the bursa copulatrix in the sample from Selcë where the junction of the uterus gland is far from the connection between the distal oviduct and the pedunculus of the bursa copulatrix in the sample from Zupanje Selo.

Figures

Fig. 105. Distribution of samples with undetermined position in the phylogenetic trees: white= Cochlostoma (T.) macei (Bourguignat, 1870); purple =C. (T.) sturanii (Wagner, 1897); orange=C. (T.) kleciaki (Braun, 1887); yellow=C. (T.) arnautorum (Wagner, 1906); red =C. (T.) atlanticum (Bourguignat, 1868); green =C. (T.) euboicum (Westerlund, 1885); blue =C. (T.) martensianum (Möllendorff, 1873); magenta= C. (T.) simrothi (Caziot, 1908); cyan =C. (T.) fontqueri Haas, 1924.