Cochlostoma (T.) pageti Klemm, 1962

Zallot, Enrico, Kamchev, Panche, Schilthuizen, Menno, Fehér, Zoltán, Mattia, Willy De & Gittenberger, Edmund, 2024, Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae), European Journal of Taxonomy 927, pp. 1-163 : 38-42

Cochlostoma (T.) pageti Klemm, 1962

Figs 26 (red dots), 27D, 32–33

Cochlostoma pageti Klemm, 1962: 213 , pl. 1 fig. 1.

Cochlostoma gracile subaiorum Schütt, 1977: 21 , pl. 1 fig. 7.

Holotype

GREECE • probably ♀; “ Paraskevi , 1500 m, im Epirus ”; May 1933; Beier leg.; NHMW74159 .

Other specimens

GREECE • 2- Terovo ; 39.4488° N, 20.8833° E; 1975; Subai leg.; NHMWK69266 • 3- Katafito ; 39.6333° N, 21.2242° E; 2011; Kontschán, Murányi, Szederjesi and Ujvári leg.; HNHM99329 .

Type locality

GREECE • 1- Epirus, Paraskevi ; 39.5682° N, 21.2607° E.

Description

SHELL. Spaced riblets on last part of protoconch. Teleoconch spotless, with prominent and rather thin ribs becoming weaker toward aperture. Moderately strong lip with indented columellar lobe covering umbilicus.

MEASUREMENTS. 5 ♀♀: whorls=7.1–8, H = 5.4–7.4 mm, H/W=2.43–2.62, roundness =0.13–0.16, ribs incl.= 56–66°, apert. incl.= 14–26°, ribs/mm 1 st wh.= 6–10, ribs/mm 4 th wh.=7–10.

FEMALE GENITAL ORGANS. Ventral-posterior connection of pedunculus to bursa copulatrix. Short, oval seminal receptacle confined to ventral side of body with 2–3 loops close to apex. Junction of uterus gland moderately far from connection between pedunculus and distal oviduct.

Remarks

The analysis of the type series indicates that C. (T.) pageti and C. gracile subaiorum Schütt, 1977 are conspecific. The short distance between the type localities supports this conclusion. In Wagner’s collection (MIZPAS -W7085) there is a specimen ( Fig. 34 ), named “pindicus ” from the Acarnania region, roughly 100 km south of the range of C. (T.) pageti . Despite the low resolution of the picture, the deep indentation of the columellar lobe is clearly visible, like in C. (T.) pageti . It remains unclear whether this population is conspecific with or only related to C. (T.) pageti .

Additional note to clade A

A terminal branch of clade A includes samples from the central and southern Apennines that can be referred to C. (T.) cassiniacum and C. (T.) adamii . They share the female genital morphology and some shell features, like the umbilicus hidden by the curved columellar lobe and the pronounced ribbing, with rounded, whitish ribs.

Figures

Fig. 26. Distribution of taxa part of clade A: cyan dots =Cochlostoma (T.) reitteri (Boettger, 1880); green dots =C. (T.) pallgergelyi sp. nov.; red dots =C. (T.) pageti Klemm, 1962. Black center indicates samples with amplified 16S.

Fig. 34. Specimen in Wagner’s collection (MIZPAS-W7085) from the Acarnania region of Greece, named as Pomatias pindicus (nomen nudum).