Pachybrachis nigricornis

Barney, Robert J., 2016, Definition and Revision of theOthonusSpecies-Group of North AmericanPachybrachisChevrolat (Coleoptera: Chrysomelidae: Cryptocephalinae), The Coleopterists Bulletin 70 (3), pp. 569-605: 569-605

publication ID

http://doi.org/ 10.1649/0010-065X-70.3.569

persistent identifier

http://treatment.plazi.org/id/03A3AA14-CB5E-3509-FC94-A3FD7ADB9630

treatment provided by

Diego

scientific name

Pachybrachis nigricornis
status

 

Pachybrachis nigricornis  

Specimens Examined. Holotypes + 1,458 specimens. CANADA: ALBERTA: Edmonton, 19.vii.1928, F. S. Carr [1♂ (Y), EUB nigricornis, FSCA   ]; Waterton Park, 16.vi.1952, L. A. Konotopetz [1♀, EUB difficilis, NDSIRC   ]. BRITISH COLUMBIA: Midday Valley, Merritt, 7.vi.1924, K. F. Auden [1♂, MCZ; 2♂♂ 2♀♀, EUB difficilis, INHS   ]. ONTARIO: Hasting Co., Baptiste Lake , 10. vii.1995, B. F. & J. L. Carr [1♂ (Y), CNC]; Muskoka Co. , Bala , 24.vi.1956, W.J. Brown [1♂ 1♀, EUB difficilis, CNC   ]; Ottawa Co. , Dirleton , 4.vii.1956, S. D. Hicks [1♂ 1♀, CNC]; Prince Edward Co. , 18.vi.1947, J. F. Brimley [2♀♀, EUB carbonarius, NDSIRC   ; 2♂♂ 1♀, EUB carbonarius, SDSU   ]; same data, except 10-17.vi.1953 [4♀♀, EUB carbonarius, NDSIRC   ; 1♂, EUB carbonarius, SDSU   ]; same data, except 23.vi.1954, wild strawberry [1♂, EUB carbonarius, NDSIRC   ; 1♂, EUB carbonarius, SDSU   ]. MANITOBA: Unknown Co. , Aweme , 7.viii.1902, E. Criddle [1♀, litigiosus, MCZ-BOW   ]; same data, except 21.vi.1904 [1♀, nigricornis, MCZ-BOW   ]; same data, except 29.vi.1906 [1♂, nigricornis, MCZ-BOW   ]; same data, except 17.vii.1907 [1♂ 1♀, nigricornis   , MCZ- BOW]; same data, except vi.1910, [1♂, HCF wahsatchensis, MCZ-FALL   ]; Mafeking , 3.vii.1937, H. T. Peters [1♀ (Y), EUB nigricornis, SEMC   ]; Winnipeg , Assinabone Park 12.vii.1984 [1♀, EUB nigricornis, FSCA   ]. USA: ALABAMA: Cleburne Co. , Cheaha State Park , 3.v.1964, taken on Senecio, E. U. Balsbaugh, Jr.   [1♀ (Y), EUB nigricornis, AUEM   ]; DeKalb Co. , DeSoto State Park , 18.v.1963, E. U. Balsbaugh, Jr. [1♀, EUB carbonarius, AUEM   ]; Houston Co. , Madrid , 18.vi.1954, D. L. Haynes [1♀, MSUC]; Lee Co. , 3 mi N of Auburn, fish ponds, 5.v.1964, ex. Quercus, E. U. Balsbaugh, Jr.   [2♀♀, EUB carbonarius, AUEM   ; 1♀, EUB carbonarius, NDSIRC   ]; Mobile Co. , Mobile , vii.1909, H. P. Loding [1♂, MCZ-FALL]; same data, except iv.1910 [1♂, MCZ-FALL]; same data, except vi.1910 [1♂, MCZ-FALL]. ARKANSAS: Baxtor Co. , Bull Shoals State Park , 3.vi.1998, D. C. Arnold [1♀ (Y), OSEC]; Hempstead Co. , Hope , 22.v.1925 [1♂, MCZ-FALL]; Logan Co. , Magazine Mountain , 15-16.vii.1949, Stannard & Sanderson [1♂, RJB autolycus, INHS   ]; same data, except 23-27.vi.1988, D. G. LeDoux [1♀, autolycus, TAMU   ]; same data, except 1350′, 17-19.v.1989, J. MacGown & T. Schiefer [2♂♂ 2♀♀, EGR autolycus, MEM   ]; same data, except 2640′, 15-20.v.1989, T. Sciefer & P. Porter [1♂ 2♀♀ (1♀ Y), EGR carbonarius, MEM   ]; Montgomery Co. , Hickory Nut Mountain , ex. Quercus sp.   , 24.vi.2002, C. L. Staines [2♂♂ 3♀♀, RJB autolycus, USNM   ]; Polk Co. , S of Board Camp R29W, T3S, Sec. 22, 6.vi.1986, C. B. Barr [1♂, EGR carbonarius, LSAM   ]; Unknown Co. , Ark [1♀, EUB carbonarius, INHS   ]. COLORADO: Archuleta Co. , Pagosa Springs , about 37°16′N, 107°0′W, 7200′ [1♀, AMNH]; Boulder Co. , Boulder 5800′, 5.vi.1961, B. H. Poole [1♂, EUB nigricornis, NEOPARATYPE, SDSU   ]; same data, except 5500′, 9.vi.1961 [2♂♂, EUB nigricornis, RJBC   ]; Costilla Co. , Garland , 19.vi, Hubbard & Schwarz [1♀ (Y), MCZ-FALL]; Douglas Co. , Pike National Forest Flat River Campground, 6.vii.1995, R. L. Blinn [1♂, NCSU]; Elbert Co. , Cedar Point , 39°21.3′N 103°52.0′W, 5800 ft., 16.vi.2012, S. M. Clark [12♂♂ 10♀♀, RJB autolycus, BYUC   ]; El Paso Co. , Colorado Springs 6,000 - 7,000 ft., 15-30.vi.1896, H. F. Wickham [1♂, HCF autolycus, MCZ-FALL   ; 2♂♂ 2♀♀, AMNH]; Jefferson Co. , Bear Creek Lake Park, Soda Lake (W of Denver), 1.vi.1985, M. F. Whiting [6♂♂, SMC nigricornis   , 1♂, SMC autolycus, BYUC   ]; Green Mountain Park, Lakewood, 6,300 ft., 24.v.1988, R. S. Beal [2♀♀ (Y), BYUC]; Park Co. , Kenosha Pass, 10,000′, 3.viii.2012, 39.4137°N 105.7547°W, R. L. Blinn [2♂♂, NCSU]; Washington Co. , Rt. 71, Last Chance, 21.vi.2009, J. Huether & R. Turnbow [1♂, RJB autolycus, BYUC   ]; Unknown Co. , [1♀, CEUM]; Colo., C. F. B. [1♀ (Y), MCZ-FALL]; Rocky Mountain National Park , 5.viii.1947, R. H. Beamer [1♂, EUB difficilis, SEMC   ]; same data, except Cascade Lodge, 2.viii.1951, H. C. Severin [1♂, EUB difficilis, SDSU   ]. DISTRICT OF COLUMBIA: Washington DC, Hubbard & Schwarz [1♀, UMRM]. FLORIDA: Nassau Co. , Amelia, 15.vi.1954, R. L. Fischer [1♀, MSUC]; Unknown Co. , Fla [1♂ 1♀, EUB carbonarius, INHS   ]. GEORGIA: Banks Co. , Lula, 30.v.1937, P. W. Fattig [1♂, UGCA]; Chatham Co. , Tybee Island , 25.vi, H.W. Wenzel [4♂♂, OSUC]; Clarke Co. , 3.vi.1940, H. O. Lund [1♀, UGCA]; Athens, 9.v.1937 [1♀, UGCA]; same data, except 22.v.1937, P. W. Fattig [1♀, UGCA]; Whitehall Forest , 5-16.v.1988, Malaise trap, C. L. Smith [4♂♂, UGCA]; same data, except 26-30.v.1988 [1♀, UGCA]; same data, except 17-23.vi.1988 [1♀, UGCA]; Cobb Co. , Kennesaw Mt. , 21.v.1939, P. W. Fattig [1♂, EUB carbonarius, NDSIRC   ]; Gwinnett Co. , Dacula, 23.v.1937, P. W. Fattig [2♀♀, UGCA]; Stone Mt. , 25.v.1928, P. W. Fattig [1♀, UGCA]; Houston Co. , Perry, 14-30.iv.1938, P. W. Fattig [6♀♀, EUB carbonarius, UGCA   ]; Jackson Co. , vicinity Attica Chr., 24.iv.1986, R. Morris [1♀, UGCA]; Jefferson Co. , Louisville, 8.v.1937, P. W. Fattig [1♀, UGCA]; Upson Co. , Thomaston, 18.v.1937, P. W. Fattig [1♀, UGCA]; Unknown Co. , Geo., Morr. [4♂♂ 1♀, MSUC]; Ga. [1?, carbonarius, MCZ-LEC   ]. IDAHO: Clark Co. , Kilgore, Ching Creek , 2.vii.2004, ex. Salix sp.   , A. C. Williams [1♀, BYUC]; Fremont Co. , Blue Creek Reservoir, 23.vii.2002, S. M. & B. D. Clark [1♀, BYUC]; Warm River , above campground, 3.viii.1982, Baumann-Winget [1♀, BYUC]; Moscow Co. , Moscow Mts. , 4.vii.1950, N. M. Downie [1♂, EUB difficilis, FMNH   ]. ILLINOIS: Calhoun Co. , Cap. Au Gres Hill Pr. , 25.v.1967, M. W. Sanderson [3♂♂ 2♀♀, RJB carbonarius, INHS   ; 1♀, RJBC]; Clendenny Hill Prairie , 2.5 mi N of Belleview, R. J. Barney [1♀ (Y), RJBC]; Kampsville, 8.vi.1949, Sanderson & Stannard [1♂ 1♀, EUB autolycus, INHS   ]; Kampsville, Perrin’ s Ledge, 15.v.1968, M. W. Sanderson [1♂, INHS]; Swarnes Hill Prairie , 1 mi N of Hamburg, 22.vi.1982, R. J. Barney [1♂ 2♀♀, RJB autolycus, RJBC   ]; Christian Co. , NE of Clarksdale, 23.vi.1961, ex. Rhus glabra, M. W. Sanderson   [1♂, INHS; 1♀, RJBC]; Clark Co. , Clarksville, 24.vi.1950, Sanderson & Benjamin [1♀, INHS]; Coles Co. , Charleston, 24.v.1970, M. A. Goodrich [1♀, EIUC]; Diona, 13.vi.1939 [1♂ 1♀, EIUC]; Effingham Co. , 1 mi S of Mason, 22.v.1963, M. W. Sanderson [3♂♂ 3♀♀, EUB carbonarius, INHS   ; 1♀, RJBC]; Hwy 37, 1 mi S Mason @ church, railroad prairie, 3.vii.2013, 38.938050N 88.643173W, R. J. & S. Barney [1♀, RJBC]; Franklin Co. , Sesser, 21.vi.1950, Sanderson & Smith [1♂, EUB carbonarius, INHS   ]; Greene Co. , Bowman Hill Prairie , N of Hillview, 26.v.1967, M. W. Sanderson [1♂ 1♀, RJB carbonarius, INHS   ]; Cole Creek Hill Prairie , 1.7 mi S of Eldred, 24.vi.1981, R. J. Barney [2♂♂ 4♀♀ (1♀ Y), RJB carbonarius, RJBC   ]; Eldred , 7.vi.1949, Stannard & Sanderson [2♂♂, EUB carbonarius, INHS   ]; Henderson Co. , Oquawka, 8.vi.1949, Stannard & Sanderson [1♀, INHS]; Iroquois Co. , Watseka, 4.vi.1932, Frison & Mohr [1♂, INHS]; Jackson Co. , Carbondale, 21.v.1908 [1♀, INHS]; same data, except 15.v.1910 [2♂♂, INHS]; Jefferson Co. , Mount Vernon , 18.vi.1993, S. M. Clark [1♀, BYUC]; Jersey Co. , Pere Marquette State Park , 17.v.1977, W. G. Ruesink [1♂ 1♀, INHS]; same data, except Ravine Trail Hill Prairie , 22.vii.1981, R. J. Barney [1♀, RJBC]; Richwood Township , 23.v.1950, Sanderson & Stannard [7♀♀ (4♀♀ Y), RJB carbonarius, INHS   ]; Johnson Co. , Goreville, 13.vi.1958, R. L. Fischer [1♂, MSUC]; Ozark, 18.v.1932, H. L. Dozier [1♂, EUB carbonarius, INHS   ]; Parker, 18.v.1908 [1♂, EUB carbonarius, INHS   ]; Kankakee Co. , 2 mi E of Wichert, 3.vi.1976, J. K. Bouseman, W. G. Ruesink & R. J. Barney [1♂ 3♀♀, EUB carbonarius, RJBC   ; 1♂, INHS]; Marion Co. , Tonti Railroad Prairie , 22.v.1963, M. W. Sanderson [2♂♂ 4♀♀, EUB carbonarius, INHS   ; 3♀♀, RJBC]; Mason Co. , Revis Springs Hill Prairie , 10.vi.1982, R. J. Barney [21♂♂ 11♀♀, RJB autolycus, RJBC   ; 1♀ (Y), RJBC]; McHenry Co. , Algonquin, W. A. Nason [1♂, MCZ-FALL; 2♀♀, INHS]; Monroe Co. , Bluff Cemetery, 1 mi E of Fountain, 26.v.1978, R. J. Barney [2♂♂, RJBC]; same data, except 12.v.1981, R. J. Barney [3♂♂ 7♀♀ (5♀♀ Y), RJB carbonarius, RJBC   ]; same data, except 7.vii.1981 [1♀ (Y), RJBC]; Fountain Bluff, 15.v.1932, Frison, Ross & Mohr [1♂ 3♀♀, EUB carbonarius, INHS   ]; Fults Hill Prairie , 4 mi SE of Fults, 12.v.1981, R. J. Barney [2♂♂ 3♀♀, RJB autolycus   + 2♂♂ 7♀♀ (4♀♀ Y), RJB carbonarius RJBC   ; 2♂♂ 3♀♀, RJB autolycus, NDSIRC   ]; same data, except 17.vi.1981 [2♂♂ 5♀♀, RJB autolycus   + 3♂♂ 7♀♀ (2♀♀ Y), RJB carbonarius, RJBC   ]; same data, except 7.vii.1981 [4♀♀, RJB autolycus   + 2♀♀, RJB carbonarius, RJBC   ]; same data, except 12.v.1982 [2♂♂, RJB autolycus   + 1♂, RJB carbonarius, RJBC   ]; same data, except 1.vii.2013, 38.1544°N 90.1848°W, R. J. & S. Barney [31♂♂ 33♀♀, RJB autolycus   + 21♂♂ 12♀♀ (1♀ Y), RJB carbonarius, RJBC   ]; Valmeyer, 24.v.1950, ex. Desmodium, M. W. Sanderson   [39♂♂ 7♀♀, EUB autolycus, INHS   ; 1♂ 1♀, RJB autolycus, RJBC   ; 1♀, carbonarius, INHS   ]; Morgan Co. , Northeast Meredosia Hill Prairie , 4.8 mi W of Arenzville, 22.vii.1981, R. J. Barney [1♂, RJB autolycus, RJBC   ]; same data, except 2.vii.2013, 39.8566°N 90.4644°W, R. J. & S. Barney [22♂♂ 10♀♀, RJB autolycus, RJBC   ]; Pulaski Co. , Pulaski, 25.v.1908 [1♀, USNM]; Rock Island Co. , 2 mi N of Cordova, 14.vii.1982, R. J. Barney [1♂, RJBC]; Whiteside Co. , Thomas-Fulton Sand Prairie , 15.vii.1986, R. J. Barney [2♀♀, RJBC]; Will Co. , Braidwood Dunes and Savanna Nature Preserve, 4.vii.2013, 41.259162N 88.195902W, R. J. & S. Barney [1♀, RJBC]; Unknown Co. , Ill. [1♀, INHS; 1?, carbonarius, MCZ-LEC   ]; same data, except H. Soltau [2♂♂ 1♀, Fall 1915 Revis, USNM]; Southern Ill. , H. Soltau [3♀♀, Fall 1915 Revis, USNM]; S ILL [4♀♀ (1Y), UMRM]. INDIANA: Clark Co. , Clarke Co. State Forest , 27.vi.1937 [1♂ 2♀♀, PERC]; Floyd Co. , 8.vi.1965 [2♀♀, EUB carbonarius, PERC   ]; same data, except 23.vi.1903, W. S. Blatchley [1♀ (Y), P. litigiosus, PERC   ]; same data, except 14.v.1965, E. Saugstad [2♂♂, EUB, NDSIRC]; Lake Co. , 26.vi.1907, W. S. Blatchley [1♀, PERC]; Lawrence Co. , 11.vi.1908, W. S. Blatchley [1♂, EUB carbonarius, PERC   ]; Orange Co. , French Lick, 28.v.1909 [2♂♂ 1♀, HCF carbonarius, MCZ-FALL   ]; Parke Co. , Turkey Run, 20.vi.1926, K. F. Auden [1♀, INHS]; Tippecanoe Co. , 27.vi.1956, N. M. Downie [2♂♂, AMNH]; same data, except 15.vi.1957 [2♀♀, AMNH]. IOWA: Unknown Co. , Ia. [2♂♂ 1♀, EUB carbonarius, ISUI   ]. KANSAS: Barber Co. , 15 mi W Medicine Lodge, 13.v.1966, Michener & party [1♂ 2♀♀, EUB autolycus, SEMC   ]; Bourbon Co. , 800 ft., R. H. Beamer [3♂♂, RJB autolycus, SEMC   ]; Elk Creek Lake , west side, 30.v.2008, D. J. Bennett [3♂♂ 9♀♀, RJB autolycus, SEMC   ]; Cherokee Co. , 2 mi S Galena, 29.v.2012, G. A. Salibury [1♂ 1♀, SEMC]; Clark Co. , 1962 ft., vi, F. H. Snow [1♀, HCF, SEMC]; Cowley Co. , 1114 ft., R. H. Beamer [1♂ 1♀, EUB autolycus, SEMC   ]; Decatur Co. , 9.vii.1932, R. H. Beamer [1♀, EUB autolycus, SEMC   ]; Hamilton Co. , 3850 ft., F. H. Snow [1♂ 1♀, HCF autolycus, SEMC   ]; Labette Co. , Big Hill Reservoir, Intersection 19000 & Douglas, 23.v.2008, A. C. Warden [1♀, SEMC]; Linn Co. , R. H. Beamer [3♀♀, EUB autolycus, SEMC   ]; McPherson Co. , 27.vii.1927, R. H. Beamer [1♂ 2♀♀, EUB autolycus, SEMC   ]; Meade Co. , Meade, 12.vi.1903 [4♂♂, HCF autolycus   , PARATYPES, MCZ-FALL]; Miami Co. , R. H. Beamer [3♂♂ 2♀♀, EUB autolycus   + 1♀, EUB carbonarius, SEMC   ]; Montgomery Co. , Elk City, 25.iv.1940, R. H. Beamer [1♂, EUB autolycus, SEMC   ]; same data, except 24.v.1940 [1♂ 1♀, EUB autolycus   + 2♀♀, EUB carbonarius, SEMC   ]; same data, except 3.vi.1940, Beamer & Sanderson [2♀♀, EUB autolycus   + 7♀♀, EUB carbonarius, SEMC   ]; Neosho Co. , Chanute, 30.viii.1933 [9♂♂ 7♀♀, EUB autolycus, SEMC   ]; St. Paul , 9 mi S Neosho Wildlife Area, 30.v.2008, A. C. Warden [2♂♂ 7♀♀, RJB autolycus, SEMC   ]; Ottawa Co. , 5 mi N Bennington, 18.vi.1954, C. D. Michener [1♂, EUB autolycus, SEMC   ]; Reno Co. , Medora , 13.vi.1915, W. Knaus [1♂, OSUC]; same data, except 8.vii.1927, R. H. Beamer [1♂ 1♀, EUB autolycus, SEMC   ]; same data, except 4.vii.1931, P. A. Readio [1♀, EUB autolycus, SEMC   ]; same data, except 25.vi.1940, D. R. Lindsay [2♀♀, EUB autolycus   + 1♀, EUB carbonarius, SEMC   ]; Sandhills State Park , 2.5 mi S Medora, 38.116°N 97.855°W, 4.vi.2007, S. M. Clark [1♀, RJB autolycus, BYUC   ; 1♀ (Y), RJB carbonarius, RJBC   ]; Rice Co. , 10.5 mi NW Medora, 4.vi.2007, S. M. Clark [1♂, BYUC; 1♀ (Y), RJBC]; Riley Co. , 7-19.vi.1962, J. R. McCoy [1♂ 1♀, EGR autolycus, MEM   ]; same data, except 8.vii.1962 [2♀♀, EGR autolycus, MEM   ]; same data, except 1-12-vi.1963 [9♂♂ 9♀♀, EGR autolycus, MEM   ]; same data, except 12.vi.1963, J. R. McCoy [2♂♂, autolycus, TAMU   ]; Rush Co. , 28.vi.1916, F. X. Williams [1♀, EUB autolycus, SEMC   ]; Saline Co. , 15.vii.1927, L. C. Woodruff [1♀, EUB autolycus, SEMC   ]; Scott Co. , 21.vi.1929, H. O. Deay [1♀, EUB autolycus, SEMC   ]; Sedgwick Co. , Wichita, 10.vi.1940, Beamer, Penner, Sanderson [1♂ 1♀, EUB carbonarius, SEMC   ]; Stafford Co. , Salt Marsh, 21.vi.1940, R. H. Beamer [1♀, EUB autolycus, SEMC   ]; Wabaunsee Co. , 17.vi.1963, J. R. McCoy [1♂, EGR autolycus, MEM   ]; Wilson Co. , 3.8 mi E Fredonia, Hwy. 17, 37.5302°N, 95.7361°W, 3.vi.2007, E. G. Riley [9♂♂ 12♀♀ (2♀♀ Y), autolycus, TAMU   ]; Woodson Co. , Biscuit Mound, 37.8651°N, 95°9284W, 3.vi.2007, E. G. Riley [5♀♀, autolycus, TAMU   ]; Unknown Co. , Kan. [2♀♀, CEUM]; same data, except Snow [1♂, HCF autolycus, SEMC   ]; Ks., Stromberg Collection [2♂♂, autolycus, INHS   ]; Kans., Snow [4♀♀, HCF autolycus, MCZ-FALL   ; 1♂, EUB autolycus, NDSIRC   ]. KENTUCKY: Christian Co. , Fort Campbell , 5.vi.2008, R. J. Barney [8♂♂ 3♀♀, carbonarius, RJBC   ]; same data, except 21.vi.2009 [1♀, RJBC]; Graves Co. , Mayfield, 20.vi.1958, J. G. Brady [1♀, MSUC]; Grayson Co. , 2 mi W of Leitchfield, 11.vi.1983, R. J. Barney [5♂♂ 5♀♀ (2♀♀ Y), carbonarius, RJBC   ]; same data, except 29.vi.1985 [3♀♀ (2♀♀ Y), carbonarius, RJBC   ]; same data, except 30.v.2006, R. J. Barney [1♀, RJBC]; Hardin Co. , Eastview Barrens State Nature Preserve, 21.vi.2004, R. J. Barney [1♀, RJBC]; same data, except 30.v.2006 [2♂♂ 1♀, RJBC]; same data, except 12.vi.2007 [1♀, RJBC]; same data, 21.v.2008 [2♀♀, RJBC]; same data, except 13.vi.2008 [1♂ 1♀, RJBC]; same data, except 9.vi.2009 [3♂♂ 3♀♀ (2♀♀ Y), carbonarius, RJBC   ]; same data, except 1.vii.2009 [1♂ 1♀, RJBC]; Hart Co. , 1 mi S of Rowletts, 29.vi.1985, R. J. Barney [1♂ 2♀♀ (1♀ Y), carbonarius, RJBC   ]; Henry Co. , KSU Environmental Education Center, 26.v.2006, R. J. Barney [4♂♂ 7♀♀, RJBC]; same data, except 13.vi.2006 [1♂, RJBC]; same data, except 19.vi.2007 [2♂♂ 1♀, RJBC]; same data, except 2.vii.2007 [1♀, RJBC]; same data, except 13.v.2008, ex. Desmodium paniculatum   [3♂♂ 3♀♀, carbonarius, RJBC   ]; LaRue Co. , Thompson Creek Glades State Nature Preserve, 30.v.2006, R. J. Barney [1♂ 1♀, carbonarius, RJBC   ]; Lewis Co. , Crooked Creek Barrens State Nature Preserve, 24.v.2006, R. J. Barney [2♂♂ 1♀, carbonarius, RJBC   ]; same data, except 17.vi.2009, R. J. Barney & S. L. Hall [2♂♂ 1♀, RJBC]; Lincoln Co. , Bouteloua Barrens State Nature Preserve, 6.vi.2005, R. J. Barney [2♂♂, RJBC]; Logan Co. , Raymond Athey Barrens State Nature Preserve, 18.v.2005, R. J. Barney [2♀♀, RJBC]; same data, except 3.vi.2005 [4♂♂ 2♀♀, RJBC]; same data, except 16.vi.2005 [3♂♂ 2♀♀, RJBC]; same data, except 9.v.2006 [4♂♂ 12♀♀ (5♀♀ Y), carbonarius, RJBC   ]; same data, except 8.vi.2006 [10♂♂ 11♀♀ (4♀♀ Y), carbonarius, RJBC   ]; same data, except 23.v.2007 [2♂♂ 3♀♀ (1♀ Y), carbonarius, RJBC   ]; same data, except 18.v.2008 [9♂♂ 13♀♀ (10♀♀ Y), carbonarius, RJBC   ]; same data, except 19.vi.2008 [5♂♂ 5♀♀, RJBC]; same data, except 19.v.2009 [3♂♂ 14♀♀ (7♀♀ Y), carbonarius, RJBC   ]; same data, except 22.vi.2009 [1♀ (Y), RJBC]; Nicholas Co. , Blue Licks Battlefield State Nature Preserve, 5.vi.1983, R. J. Barney [1♀, RJBC]; Pendleton Co. , Kincaid Lake State Park , 30.vi.1971, P. H. Freytag [1♂, UKIC]; Robertson Co. , Blue Licks Battlefield State Nature Preserve, 24.v.2006, R. J. Barney [3♂♂ 4♀♀, carbonarius, RJBC   ]; same data, except 15.vi.2006 [2♀♀, RJBC]; same data, except 20.vii.2006 [1♀, RJBC]; same data, except 22.v.2008 [1♂ 1♀, RJBC]; same data, except 17.vi.2009, R. J. Barney & S. L. Hall [1♂ 5♀♀, RJBC]; same data, except 30.vi.2009 [1♂ 3♀♀, RJBC]; same data, except 24.vii.2009 [2♀♀, RJBC]; Trigg Co. , Fort Campbell , 5.vi.2008, R. J. Barney [10♂♂ 4♀♀, RJBC]; same data, except 18.vi.2008 [7♂♂ 15♀♀ (4♀♀ Y), carbonarius, RJBC   ]; same data, except 10.vii.2008 [1♂ 2♀♀, RJBC]; same data, except 18.v.2009 [1♂, RJBC]; same data, except 1.vi.2009 [12♂♂ 13♀♀ (2♀♀ Y), carbonarius, RJBC   ]; same data, except 21.vi.2009 [6♂♂ 3♀♀ (1♀ Y), carbonarius, RJBC   ]. LOUISIANA: Bossier Par., Barksdale AFB, 22-30.iv.1996, Malaise trap in prairie, D. M. Pollock [1♀ (Y), MEM]; Barksdale Air Force Base, 32° 29′N, 93° 35′W, 19.v.1996, night sweep, calcareous prairie, E. G. Riley [3♂♂ (1♂ Y) 26♀♀ (23♀♀ Y), TAMU]; Bodcau W.M.A. 32° 42′N 93°30′W, 23.v.2000, V. L. Moseley [1♀, LSAM]; Bodcau Wildlife Man. Area , 32° 42′N, 93 o 30′W, 20.v.1996, E. G. Riley [1♂, TAMU]; Lamar Co. , Camp Maxey, 33°48′36′N, 95°34′10′W, 24.v.2003, E. G. Riley [8♂♂ 7♀♀, TAMU]; same data, except [3♂♂ 10♀♀, autolycus, TAMU   ]; Natchitoches Pr., Kisatchie Natl. Forest ca. 2 mi E Red Bluff Campground, 9.iv.1992, D. A. Rider [2♀♀, LSAM]; same data, except 7.v.1993 [1♀, LSAM]; Rapides Pr., Valentine Lake near Gardner, 6.v.1972, M. E. Dakin & P. L. Dakin [1♀, LSAM]. MAINE: Androscoggin Co. , Wales, 19.vi.1906, C. A. Frost [1♂, HCF difficilis, MCZ-FALL   ]; Hancock Co. , Mt. Desert , 16.vi.1922, C. W. Johnson [1♂, difficilis, USNM   ]; Kennebec Co. , Augusta, 6.vii.1940, A. E. Brower [2♂♂, difficilis, USNM   ]. MARYLAND: Anne Arundel Co. , Harmans, 10.vi.1985, J. F. Cavey [1♀, JFCC]; same data, except 2.vii.1985 [1♀, JFCC]; Howard Co. , Ellicott City, 4.vi.1968, R. Gordon [1♀, USNM]; Prince George’ s Co., 9.vi.1960, J. R. McCoy [1♂, MEM]; College Park, 2.v.1948, B. K. Dozier [1♀, FSCA]. MASSACHUSETTS: Barnstable Co. , Wellfleet, 25.vi.1920 [1♀, MCZ-FALL]; same data, except 27.viii.1901 [1♀, MCZ]; Middlesex Co. , Sherbon, 15.vi.1913, C. A. Frost [2♂♂ 2♀♀, difficilis MCZ   ]; Tyngsboro, 15.vii.1900 [1♂, HCF difficilis   , PARATYPE, MCZ-FALL]; Unknown Co. , Mass. [1♀, MCZ-FALL; 2??, difficilis, MCZ-LEC   ]. MICHIGAN: Allegan Co. , 10.vi.1949, R. Namba [1♂, MSUC]; Cheboygan Co. , 3.v.1930 [1♀, EUB difficilis, SEMC   ]; Douglas Lake , 28.vii.1926, C. Martin [1♀, EUB difficilis, SEMC   ]; Chippewa Co. , Whitefish Point, 28.vii.1914, A. W. Andrews [1♀, EUB difficilis, MSUC   ]; Kalamazoo Co. , Gull Lake Biological Station , 5.vi.1957, R. L. Fischer [1♂, MSUC]; same data, except 29.vi.1964 [1♀, MSUC]; same data, except 26.v.1976, R. J. Barney [1♀ (Y), carbonarius, RJBC   ]; Livingston Co. , Whitmore Lake , 7.vii.1956, G. H. Nelson [2♂♂ 2♀♀, AMNH]; Marquette Co. , Marquette, 7.viii, Hubbard & Schwarz [3♀♀ (2♀♀ Y), Fall 1915 Revis carbonarius, USNM   ]; Montcalm Co. , Stanton, 5.vi.1959, H. D. Niemczyk [1♀, MSUC]; Newaygo Co. , Newaygo, 30.v.1928, L. G. Gentner [1♀, MSUC]; Wayne Co. , Detroit, Hubbard & Schwarz [2♀♀, Fall 1915 Revis difficilis, USNM   ]; Unknown Co. , MI [1♀, teste Fall, CMNH]; MI, Lec. [1♀, CMNH]. MINNESOTA: Anoka / Isanti Co. , Cedar Creek Natural History Area , 15.vi.2002 [1♀, UMSP]; Isanti Co. , Cedar Creek Natural History Area , 16.vii.1985 [1♂ (Y), UMSP]; same data, except 11.vi.1987 [1♂ (Y), UMSP]; same data, except 20.vi.2001 [1♂, UMSP]; Winona Co. , Winona, 2.vii.1968, Harris & Colley [1♂, UMSP]. MISSISSIPPI: Chickasaw Co. , Buena Vista , 33°53′40′N 88°49′22′W, 7.v.2010, J. G. Hill [1♂ 2♀♀ (1♂ Y), MEM]; same data, except Pulliam Prairie , 33.89416°N, 88.82416°W, 6.vi.2014, R. J. Barney [6♂♂ (1♂ Y) 4♀♀ (1♀ Y), carbonarius, RJBC   ]; Tombigbee National Forest , 5 km NNW Buena Vista, 33.9725°N, 88.8561°W, 6.vi.2014, S. M. Clark [1♀, BYUC]; Harrison Co. , Cat Island , 7.ix.1920, H. L. Dozier [1♀, SEMC]; Lowndes Co. , 11.v.1991, D. M. Pollock [1♂, MEM]; Crawford, 33°18′01′N 88°36′38′W, 27.v.2009, J. G. Hill [4♂♂ 3♀♀, MEM]; Noxubee Co. , Noxubee National Wildlife Refuge, 33°15′12′N 88°46′16′W, 11.v.2011, T. L. Schiefer [24♂♂ 20♀♀, MEM]; Oktibbeha Co. , A &M College, 17.v.1925, R. Melvin [1♂, MEM]; State College, 11.iv, C. L. Barry [1♀, MEM]; same data, except 30.iv.1939, W. L. Lewis [1♀, MEM]; Chickasaw WMA, 33.9725°N, − 88.8550°W, 6.vi.2014, R. J. Barney [1♂ 4♀♀, carbonarius, RJBC   ]; Osborn, 33°30′41′N 88°44′08′W, 13.iv.2001, J. G. Hill [1♀ (Y), MEM]; same data, except 14.v-13.vi.2001 [2♂♂ 2♀♀, MEM]; same data, except 27.v.2009 [1♀, MEM]; Osborn Prairie , 28.iv.2007, R. Turnbow [3♂♂ 1♀, FSCA]; same data, except 33.5113°N, − 88.7355°W, 6.vi.2014, R. J. Barney [1♂, RJBC]. MISSOURI: Benton Co. , Rock Hill prairie, 14.vi.1994, D. G. LeDoux [1♂ 2♀♀, TAMU]; Callaway Co. , Fulton, 24.iv.1949, G. W. Thomas [1♀, EGR carbonarius, UMRM   ]; Tucker Prairie , 4.vi.1968 [1♀, EGR carbonarius, UMRM   ]; Camden Co. , Camdenton, 11.vi.1939, N. L. Wright [1♀, UMRM]; Ha Ha Tonka State Park , Lodge Glade, 24.iv.2001, D. G. LeDoux [5♂♂ 1♀, autolycus, TAMU   ]; same data, except 17.v.2001 [18♂♂ 9♀♀ carbonarius   , 18♂♂ 8♀♀, autolycus, TAMU   ]; same data, except 21.vi.2001 [6♂♂ 10♀♀, autolycus, TAMU   ]; Rt. J NE Mack’ s Creek, 27.vi.1993, D. G. LeDoux [1♂ 2♀♀ (Y), TAMU]; Christian Co. , Sparta, 15.vi.1940, C. Wingo [1♀, UMRM]; Cooper Co. , 21.v.1939, W. R. Enns [1♀, UMRM]; Franklin Co. , Fern Glen, 14.vi.1901, G. W. Bock [2♂♂ 3♀♀, EGR autolycus, UMRM   ; 2♂♂, EUB, UMRM]; Iron Co. , Taum Sauk State Park , Wiemer Hill, igneous glade, 25.iv.2000, D. G. LeDoux [15♂♂ 32♀♀ (21♀♀ Y), TAMU]; Jackson Co. , Englewood, Devils Backbone, 17.v.1956, W. R. Enns [1♀, EGR autolycus, UMRM   ]; Jasper Co. , Wah-Sha-She Prairie , 2 mi N Asbury, 12.vi.1978, E. G. Riley [1♂ 1♀, EGR autolycus, UMRM   ]; Jefferson Co. , Pevely, 22.vi.1909, Dr. G. W. Bock [1♂ 3♀♀, EUB autolycus, UMRM   ]; Victoria Glades, 2 mi SE Hillsboro, 19.v.1995, D. G. LeDoux & B. L. LeDoux [2♂♂ 3♀♀, autolycus, TAMU   ]; Laclede Co. , Bennett Springs Savanna Nature Area , 3.vi.1995, D. G. LeDoux [1♂ 2♀♀, TAMU]; Lawrence Co. , Marionville, 4.vi.1938, W. W. Smith [1♂, EUB carbonarius, UMRM   ]; Maries Co. , Spring Creek Nature Area , 26.v.1994, D. G. LeDoux [1♂ (Y), 1♀, autolycus, TAMU   ]; Phelps Co. , Rosati, 24.v.1949, G. W. Thomas [1♂ 5♀♀ (4♀♀ Y), EGR carbonarius, UMRM   ]; Pike Co. , New Hartford, 13.vi.1940, W. S. Craig [2♀♀, UMRM]; Randolph Co. , Rothwell Park, Moberly, 14.vi.1980, E. G. Riley [1♂ 1♀, TAMU]; Reynolds Co. , Johnson Shut-Ins State Park , Horseshoe Glades, 15.vi.1995, D. G. LeDoux [2♂♂, TAMU]; St. Charles Co. , Weldon Springs, 17.vi.1968 [1♀, EGR carbonarius, UMRM   ]; St. Clair Co. , Lichen Glade Natural Area , 12.v.1994, D. G. LeDoux [1♀, TAMU]; Taberville Prairie , 14.vi.1994, D. G. LeDoux [2♀♀, TAMU]; same data, except 1.5 mi E Appleton, 10.vi.1979, E. G. Riley [1♀ (Y), EGR carbonarius, UMRM   ]; same data, except 2.5 mi N Taberville, 38.0482°N, - 93.9986°W, 2.vi.2007, R. J. Barney [5♀♀, RJBC]; St. Louis Co. , Creve Coeur Lake , 25.v.1902, G. W. Bock [2♂♂, EGR carbonarius, UMRM   ]; Jefferson, 16.vii.1975, E. G. Riley [1♀, EGR carbonarius, UMRM   ]; St. Louis , [1♂ 2♀♀, UNSM]; same data, except 3.vi.1895 [2♀♀ (Y), EUB carbonarius, UMRM   ]; same data, except 4.vi.1896 [1♀ (Y), EUB carbonarius, UMRM   ]; same data, except 9.vii.1896 [1♂, EUB carbonarius, UMRM   ]; same data, except 16.vi.1919, Dr. G. W. Bock [2♂♂, EUB, UMRM]; Stoddard Co. , 2.8 mi NE of Dexter, Holly Preserve , 10.vi.1975, E. G. Riley [1♀ (Y), EGR carbonarius, UMRM   ]; Stone Co. , 1 mi E Jct. 160 & 248, 7.vi.1975, S. E. Thewke [3♀♀ (2♀♀ Y), EGR carbonarius, UMRM   ]; Wayne Co. , 3.5 mi N of Wappapello on Rt. Z, 11.vi.1975, E. G. Riley [1♀ (Y), EGR carbonarius, UMRM   ]. MONTANA: Big Horn Co. , Bighorn River Ft. Smith, 25.v.1988, D. L. Gustafson [1♀, difficilis, BYUC   ]; Gallatin Co. , Bozemen, Gallatin Valley , 4.viii.1949, L. Ellig [2♀♀ (Y), EUB difficilis, SDSU   ]; Rosebud Co. , Highway I-94 at Reservation Creek , 18.v.1988, S. M. Clark [1♀ (Y), BYUC]; Valley Co. , Glasgow, 27.vi.1898 [1♀, MCZ-FALL]. NEBRASKA: Antelope Co. , Neligh, 18.vi.1909, W. Thompson [1♀, UNSM]; same data, except 21.vi.1909, J. T. Zimmer [1♂, UNSM]; Cumming Co. , West Point , vi.1884 [2♂♂, UNSM]; Jefferson Co. , 5-9.vi.1989, C. A. Springer [1♀, UNSM]; Keith Co. , N & E sides Lake McConaughy , 21.v.2007, J. Huether [1♂ 1♀, RJB autolycus, BYUC   ]; Lancaster Co. , Lincoln, v [1♀, UNSM]; Nine-Mile Prairie , 40°51′58.8′N, 96°48′20.1′W, elev. 397m, sweeping at night, 19.vi.2013, K. Miwa [1♀, BYUC]; Sheridan Co. , Hay Springs, 24.vi.1923 [1♀, UNSM]; Thomas Co. , Halsey, 8.vi.1911, J. T. Zimmer [1♀ (Y), UNSM]; same data, except 1.vii.1956, R. Henzlik [2♂♂, 1♂ EUB difficilis   , 1♂ EUB carbonarius, UNSM   ]; same data, except 10.vi.1958 [2♂♂ (1♂ Y), UNSM]; Unknown Co. , Sand Hills, vii [1♀, UNSM]. NEW HAMPSHIRE: Hillsborough Co. , Manchester, 15.vi [1♂, FCB carbonarius, CNC   ]; Merrimack Co. , Webster 1480, Fiske [1♂, EGR difficilis, UNH   ]; Unknown Co. , NH, [1?, litigiosus, MCZ-HORN   ]. NEW JERSEY: Burlington Co. , Woodmansie, 11.vi.1997, S. M. Clark [1♀, BYUC]; Camden Co. , Clementon, 14.v.1899, G. M. Greene [1♀, USNM]; Lucaston, 30.v.1906 [1♀, USNM]; Cape May Co. , Anglesea, 31.v, Shoemaker [1♂, USNM]; Essex Co. , Montclair, 8.vi, E. A. Bischoff [1♂, AMNH]; Gloucester Co. , Malaga, 15.vi.1912, Shoemaker [1♀, USNM]; Unknown Co. , N.J., J. B. Smith [1♂, Fall 1915 Revis, USNM]. NEW MEXICO: Cibola Co. , Mt. Taylor , C. Hoff [1♂ 2♀♀, EUB autolycus, AMNH   ]; Grant Co. , Silver City, 23.vii.1940, D. R. Lindsay [1♀, EUB autolycus, SEMC   ]; San Miguel Co. , Cowles, 18.vii.1936, M. B. Jackson [1♀, EUB difficilis, SEMC   ]; Unknown Co. , N. Mex., Dury [3♂♂, autolycus, CMNH   ; 1♂, HCF autolycus, MCZ-FALL   ; 1♂, EUB autolycus, SDSU   ]; N. M. [2♂♂, EGR autolycus, UMRM   ]; Tallque, 26.vi.1945, B. Hodgen [1♂, EUB autolycus, SEMC   ]. NEW YORK: Delaware Co. , 29.vii.1965, R. A. Moore [1♀, EUB difficilis, CUIC   ]; Nassau Co. , Massapequa L.I., Shoemaker [1♀, USNM]; Suffolk Co. , 17.vi.1967, M. I. Blenderman [1♂ 4♀♀, AMNH]; same data, except 2.vii.1967 [2♀♀, AMNH]; same data, except 10.vii.1968 [1♂, AMNH]; Unknown Co. , N. Y. City & vicinity, C. Schaeffer [3♂♂ 1♀, USNM]; Central Park L.I., 29.vi.1913, Shoemaker [1♀, USNM]; Van Courtland, 11.vii.1895, J. L. Zabriske [1♀, USNM; 1♂ 1♀, AMNH]; Mosholu [1♂, AMNH]. NORTH CAROLINA: Harnett Co. , Dunn, 17.v.1965, R. C. Froeschner [1♂, USNM]; Mecklenburg Co. , Davidson, 8.vi.1959, T. Daggy [1♀, NCSU]; Montgomery Co. , 12 mi NE Troy, 35.4978°N, 80.0357°W, 18.vi.2013, R. J. Barney [1♂ 2♀♀, carbonarius, RJBC   ]; Moore Co. , Carthage, 25.iv.1938, W. J. Gertsch [1♀, AMNH]; Southern Pines , 6.v.1915, A. H. Manee [1♀, NCSU]; same data, except 19.vi.1915 [1♀, NCSU]; same data, except 13.vi.1917 [1♀, USNM]; same data, except 20.v.1919 [1♀, EUB carbonarius, NCSU   ]; same data, except 13.v.1920 [1♂, EUB carbonarius, NCSU   ]; Wake Co. , 17.v.1962, F. L. Roberts [1♀, NCSU]; Raleigh, 21.v.1952, B. K. Dozier [2♂♂ 4♀♀, FSCA]; same data, except 23.v.1933, D. L. Vray [4♂♂ 1♀, EUB carbonarius, NCSU   ]; same data, except 27.iv.1936, C. S. Brimley [1♀, EUB carbonarius, NCSU   ]; Schenck Forest , 11.v.1994, R. L. Blinn [1♀, USNM; 1♀, NCSU]; Washington Co. , Roper, 2.v.1952, B. K. Dozier [1♂, FSCA]. NORTH DAKOTA: Barnes Co. , Euphorbia esula   L., D. A. Mundal [1♀ (Y), EUB nigricornis, NDSIRC   ]; Benson Co. , Knox, 28.vii.1937, R. H. Beamer [1♀ (Y), EUB carbonarius, SEMC   ]; Cass Co. , Fargo, 27.vi.1923, W. Baker [1♀ (Y), EUB nigricornis, NDSIRC   ]; Dunn Co. , 17.vii.1962 [1♀, EUB carbonarius, NDSIRC   ]; same data, except 23.vi.1963, R. Gordon [1♂ (Y), EUB carbonarius, NDSIRC   ]; same data, except 24.vi.1964, D. Aarhus [1♀ (Y), EUB carbonarius, NDSIRC   ]; same data, except 6.vii.1967 [1♂ (Y), EUB, NDSIRC]; Killdeer Mts. , 22.vi.1963, D. Aarhus [1♀, EUB autolycus, NDSIRC   ]; same data, except 24.vi.1964 [1♀, EUB difficilis, NDSIRC   ]; same data, except 3.viii.1965, W. Kotchman [1♀, EUB autolycus, NDSIRC   ]; 5 mi W Killdeer, 26.v.1977, E. U. Balsbaugh, Jr. [1♂, EUB nigricornis, NDSIRC   ]; Eddy Co. , 24.vii.1958, D. Aarhus [1♀ (Y), EUB nigricornis, NDSIRC   ]; Hamar, 27.vii.1937, C. L. Johnston [1♂, EUB carbonarius, SEMC   ]; McKenzie Co. , Theodore Roosevelt, 22.vi.1965, W. Kotchman [1♀, NDSIRC]; Slope Co. , Chalky Buttes, 1.vii.1965, W. Kotchman [1♂, EUB autolycus, NDSIRC   ]; Unknown Co. , Black Hills [1♂ 1♀, HCF autolycus, MCZ-FALL   ]. OHIO: Athens Co. , 28.vi.1938, J. Hughes [1♀, UGCA]. OKLAHOMA: Beaver Co. , 4 mi W Slapout, 12.vii.1971, D. C. Arnold [1♀, OSEC]; Bryan Co. , Brown, 24.v.1978, D. C. Arnold [5♀♀, OSEC]; Comanche Co. , 0.75 mi SE Medicine Park, 34.7234°N, 98.4885°W, 6.vi.2007, R. J. Barney [1♂ (Y) 1♀, RJBC]; 2 mi E Medicine Park, 34.724°N, 98.475°W, 6.vi.2007, S. M. Clark [1♀, BYUC]; Wichita Mountains near Quanah Parker Lake , 2.v.1998, S. M. Clark [5♂♂, BYUC]; Delaware Co. , Jay, 8.vii.1931, W. D. Davis [1♀, EUB difficilis, OSEC   ]; Latimer Co. , v.1986, K. Stephan [3♀♀ (Y), TAMU]; same data, except iv.1988 [1♂ (Y), TAMU]; same data, except vi.1989 [1♀ (Y), TAMU]; 5 mi W Red Oak, 14.v.1977 [1♀ (Y), TAMU]; same data, except v.1980 [1♂ (Y), TAMU]; same data, except v.1981 [2♂♂ (Y) 5♀♀ (Y), 1♂, autolycus, TAMU   ]; same data, except v.1982 [5♂♂ (Y) 7♀♀ (Y), TAMU]; same data, except v.1985 [1♂, TAMU]; same data, except v.1993 [5♀♀ (Y), TAMU]; Red Oak , 23.v.1966, D. C. Arnold [1♀, EUB difficilis   ; 1♀, EUB nigricornis, OSEC   ]; Logan Co. , 4 mi N, 1.5 mi E jct. Hwy. I-35 & St. 33, 6-12.vi.1984 [2♀♀, OSEC]; Mayes Co. , Chouteau, 30.v.1972, D. C. Arnold [1♂ 1♀, OSEC]; Locust Grove, 36°11′54′ N, 95°08′55′W, 18.v.2002, E. G. Riley [3♀♀ (Y), TAMU]; McCurtain Co. , 8.vi.1931, Costner & Davis [1♂, EUB autolycus, OSEC   ]; Noble Co. , Perry, 16.v.1972, J. Pickle [2♂♂ 1♀, OSEC]; Oklahoma Co. , Jones, 13.vi.1957, D. Alexander [1♂, EUB autolycus, OSEC   ]; Oklahoma City (Massey Farm), 14.v.1971, J. Pickle [3♂♂ 2♀♀, EUB autolycus, OSEC   ]; W of Edmond, 35°38′ 20"N; 97°28′W, 2.v.1979, K. Rudkin [1♀, OSEC]; Pawnee Co. , Sooner Lake , Pawnee, 21.vi.1984 [1♀, OSEC]; Payne Co. , 6.5 mi W, 4.5 mi SW jct. Hwy. 51 & 177, 6.vi.1984 [1♀, OSEC]; Lake Carl Blackwell , 8.vi.1959, W. A. Drew [1♂, EUB autolycus, OSEC   ]; same data, except 15.v.1979, R. White [1♀, OSEC]; nr Lake Carl Blackwell , Malaise trap, 23.v.1978, A. Hollander [1♂ 3♀♀, OSEC]; Sanborn Lake , Stillwater, 18.v.1985, D. C. Arnold [2♀♀, OSEC]; Stillwater, 9.vi.1954, F. A. Fenton [1♀, EUB autolycus, OSEC   ]; same data, except 15.v.1964, F. Miller [1♂, EUB autolycus, OSEC   ]; same data, except 2.v.1966, P. Semtner [2♂♂ 1♀, EUB autolycus, OSEC   ]; same data, except 26.vi.1969, J. Pickle [1♀, EUB autolycus, OSEC   ]; same data, except 30.vi.1971 [1♀, EUB autolycus, OSEC   ]; Pittsburg Co. , Crowder (lakeside park), 35°06′08′N, 95°41′30′W, 2.v.2004, E. G. Riley [3♂♂ 8♀♀, autolycus, TAMU   ]; Pottawatomie Co. , 10 mi S Maud, 28.v.1966, P. Semtner [1♂, EUB autolycus, OSEC   ]; Earlsboro, 26.vi.1978, D. C. Arnold [1♀ (Y), OSEC]; Rattan, 22.v.1980, D. C. Arnold [1♀ (Y), OSEC]; Stephens Co. , Clear Creek Lake , 2.v.1998, S. M. Clark & S. A. Wells [2♂♂ 2♀♀, RJB autolycus, BYUC   ]; Woods Co. , 7 mi E Waymoka, 22.vi.1972, D. C. Arnold [1♀, OSEC]; 14 mi N 9 mi W Alva, B. & J. Smith [5♂♂, autolycus, TAMU   ]; PENNSYLVANIA: Dauphin Co. , Linglestown, 21.vi, Kaeber [2♂♂, USNM]; Monroe Co. , Water Gap, C. Palm [1♀, AMNH]; Montgomery Co. , Miquon, 29.v.1960, G. B. Sleesman [1♂ 1♀, EUB carbonarius, NDSIRC   ]. SOUTH CAROLINA: Beaufort Co. , Beaufort, 28.iv [1♀, CUAC]; Beaufort & Hampton Co. , Yemassee, 14.vi.1954, R. L. Fischer [1♀, MSUC]; Charleston Co. , Meggett, 26.v.1934, O. L. Cartwright [1♂, CUAC]; Chester Co. , Fort Lawn , 13.vi.1954, R. L. Fischer [1♀, MSUC]; Chesterfield Co. , Cheraw, 28.v.1961, V. M. Kirk [1♂, EUB, NDSIRC]; Sand Hills State Forest , 26.v.2006, S. M. Clark & E. G. Riley [6♂♂ 3♀♀, BYUC]; Florence Co. , Florence, 23.iv.1935, J. E. Webb, Jr. [1♀, CUAC]; Georgetown Co. , Georgetown, 18.v.1932, O. L. Cartwright [1♀, CUAC]; Pickens Co. , Clemson College, 10.v.1932, J. G. Watts [1♂ 1♀, CUAC]; same data, except 3.v.1935 [1♂, CUAC]; Rocky Bottom, 22.v.1934, J. A. Berly [1♂ 1♀, CUAC]; Williamsburg Co. , Kingstree, 25.v.1937, J. G. Watts [2♀♀, CUAC]; Unknown Co. , Meredith, 14.v.1927, O. L. Cartwright [1♂, CUAC]; same data, except 4.v.1928 [1♀, CUAC]. SOUTH DAKOTA: Bennett Co. , Martin, 27.vi.1937, H. C. Severin [1♂, EUB nigricornis, SDSU   ]; Brown Co. , Hecla, 20.vi.1940, H. C. Severin [1♀, EUB nigricornis, SDSU   ]; Buffalo Co. , Ft . Thompson, 21.vi.1944, H. C. Severin [1♀, SDSU]; same data, except 27.vi.1946, J. A. Lofgren [1♂ (Y), EUB nigricornis, SDSU   ]; Butte Co. , Newell, 27.vi.1952, H. C. Severin [1♀, EUB difficilis, SDSU   ]; Codington Co. , Medicine Lake near Florence, 2.vi.1939, H. C. Severin [1♂, EUB nigricornis, SDSU   ]; Pot hole near Florence, 2. vi.1939, H. C. Severin [1♂, EUB nigricornis, SDSU   ]; Custer Co. , Custer, 3.vii.1932, F. R. Bingham [1♀, EUB nigricornis, SDSU   ]; Dewey Co. , Lantry, 28.vi.1947, H. C. Severin [1♀, EUB nigricornis, SDSU   ]; Hand Co. , Vayland, 7.vi.1949, H. C. Severin [1♀, EUB nigricornis, SDSU   ]; Harding Co. , Buffalo, 26.viii.1924, G. I. Gilbertson [1♂, EUB difficilis, SDSU   ]; same data, except 28.vi.1947, H. C. Severin [1♂, EUB difficilis, NDSIRC   ; 3♀♀ (Y), EUB nigricornis, SDSU   ]; Slim Buttes near Reva, 5.viii.1953, H. C. Severin [2♀♀ (Y), EUB nigricornis, SDSU   ]; Hyde Co. , Highmore, 14.vi.1948, H. C. Severin [1♂, EUB nigricornis, SDSU   ]; Jones Co. , Capa, 1.vi.1921, H. C. Severin [1♀, INHS; 1♂ (Y) 1♀ (Y), EUB nigricornis, SDSU   ]; Murdo, 43°53.1′N 100°42.4′W, 2320 ft., 3.vi.2012, S. M. Clark [1♂, BYUC]; Lawrence Co. , Hanna, Black Hills, 5.viii.1969, E. U. Balsbaugh, Jr. [1♀, EUB nigricornis, NDSIRC   ]; Lead, 13.vi.1968, V. M. Kirk [1♂ 1♀ (Y), EUB nigricornis, NDSIRC   ]; Meade Co. , Sturgis, 22.vi.1954, H. C. Severin [1♂, EUB nigricornis, NDSIRC   ; 5♂♂ (1♂ Y) 16♀♀ (12♀♀ Y), EUB nigricornis, SDSU   ]; Mellette Co. , White River , 21.vi.1931, H. C. Severin [1♀, EUB nigricornis, SDSU   ]; Mercer Co. , Lakeview, 20.vi.1930, G. I. Gilbertson [1♀, EUB nigricornis, SDSU   ]; Pennington Co. , Rapid City, 15.vi.1965, V. M. Kirk [1♂ (Y), EUB nigricornis, NDSIRC   ]; Ziebach Co. , Dupree, 25.vi.1947, H. C. Severin [1♀, EUB nigricornis, SDSU   ]; Unknown Co. , Black Hills, 11.vii.1901, J. L. Webb [1♀, MCZ-FALL]; Fox Ridge, 28.vi.1947 [1♂, EUB nigricornis, SDSU   ]. TENNESSEE: Coffee Co. , Arnold Engineering Development Center, 3.viii.1998, R. B. [1♀, UTKC]; Manchester, 21.vi.1941, W. S. Craig [1♀, UMRM]; Decatur Co. , Carroll Cabin Barren Natural Area , 35.4667°N, 87.0538°W, 8.vi.2014, S. M. Clark & E. G. Riley [1♀, BYUC]; Henderson Co. , Natchez Trace St. Forest , 8.vi.1971, D. K. Young [1♀, MSUC]; Montgomery Co. , Fort Campbell , 18.v.2009, R. J. Barney [6♂♂ 7♀♀ (1♀ Y), carbonarius, RJBC   ]; same data, except 36.588724°N, − 87.553291°W, 2.vi.2009 [4♂♂ 8♀♀, carbonarius, RJBC   ]; Rutherford Co. , Flat Rock Cedar Glades State Natural Area, 35°51′31′N 86°17′44′W, 2.vi.2010, J. G. Hill [4♂♂ 1♀, MEM]; same data, except 35.8585°N, − 86.2956°W, 4.vi.2014, R. J. Barney & S. M. Clark [2♂♂ 5♀♀, carbonarius, RJBC   ; 1♂ 1♀, BYUC]; Murfreesboro, 18.vi.2000, J. F. Cavey [1♀, JFCC]; Shelby Co. , 23.v.1948, B. Summerville [1♂, MSUC]; same data, except 10.v.1948 [2♀♀, MSUC]; Stewart Co. , Fort Campbell , 36.636903°N, − 87.648067°W, 5.vi.2008, R. J. Barney [7♂♂ 6♀♀ (1♀ Y), carbonarius, RJBC   ]; same data, except 2.vi.2009 [3♂♂ 3♀♀, RJBC]; Wilson Co. , Cedars of Lebanon State Park , 36.0767°N, − 86.3957°W, 4.vi.2014, R. J. Barney [4♂♂ 2♀♀, carbonarius, RJBC   ]; Vesta Cedar Glade State Natural Area , 36.0927°N, 86.3499°W, 3.vi.2014, R. J. Barney [1♂ 1♀, RJBC]. TEXAS: Archer Co. , Highway 281, 3 mi N Scotland, 33°42.8′N 98 o 28.7′W, 2.v.2002, S. M. Clark [4♂♂ 7♀♀, RJB autolycus, BYUC   ]; Armstrong Co. , 4 mi N Prairie Dog Town, Fork of Red River , Hwy. 207, 30.iv.2002, S. M. Clark & D. J. Cavan [1♂, BYUC]; Brazos Co. , 23.iv.1956, H. R. Burke [2♀♀, autolycus, TAMU   ]; College Station , 17-27.iv.1932, H. J. Reinhard [25♂♂ 19♀♀, autolycus   + 4♂♂ 2♀♀, EUB difficilis, TAMU   ]; same data, except 14.iv.1962, S. G. Wellso [1♀, autolycus, TAMU   ]; same data, except 24.iv.1976, J. A. Jackman [1♂, INHS]; Fannin Co. , Fannin Lake , 33°46′47′N, 96°09′34′W, 3.v.2004, E. G. Riley [15♂♂ 6♀♀, EGR autolycus, TAMU   ]; Fayette Co. , 3.v.1931, R. K. Fletcher [2♂♂ 1♀, EUB difficilis, TAMU   ]; Gray Co. , 6.5 mi W Mobeetie, 35°31′02′N, 100°33′37′W, 29.v-2.vi.2004, E. G. Riley [14♂♂ 23♀♀, EGR autolycus, TAMU   ]; Jack Co. , near Antelope, 33°23.6′N 98 o 21.9′W, 2.v.2002, S. M. Clark & D. J. Cavan [5♂♂ 1♀, RJB autolycus, BYUC   ]; Jeff Davis Co. , 5 mi NW Ft. Davis, 5000′, 5.vii.1968, J. E. Hafernik [1♂, autolycus, TAMU   ]; Lamar Co. , Gibbons Lake , 33.7002°N, 95.6506°W, 31.v.2007, R. J. Barney [1♂, RJBC]; Pat Mayse State WMA, 33.8101°N, 95.6825°W, 31.v.2007, R. J. Barney [1♂ (Y) 1♀ (Y), RJB carbonarius, RJBC   ]; Moore Co. , 1 mi E Jct. Hwy. 87/287 on FM, 35°41′25′N, 101°56′36′W, 17.v.2002, E. G. Riley [33♂♂ 41♀♀, EGR autolycus, TAMU   ]; Stonewall Co. , Jct . Hwy. 280 & Salt Fork of Brazos River , 33°10′01′N, 100°37′59′W, 11.v.2002, E. G. Riley [1♂, EGR autolycus, TAMU   ]; Wichita Co. , Burkburnett, Red River , 26.vi.1948, C. & P. Vaurie [1♀, AMNH]. UTAH: Cache Co. , 9.vi.1959, G. F. Knowlton [1♀, difficilis, BYUC   ]; Lewiston, 26.vi.1950, J. V. Bruce [1♀, EUB difficilis, MSUC   ]; Salt Lake Co. , Bingham Canyon, 16.vi.1938, D. E. Hardy & G. S. Stains [1♀, EUB difficilis, BYUC   ]; Utah Co. , Uinta National Forest , Blackhawk Campground, 26.vi.1990, S. M. Clark [1♀, difficilis, BYUC   ]; Wasatch Co. , vi.1927 [3♂♂ 6♀♀, HCF wahsatchensis   PARATYPES, “much like lustrans N. B. III p92”, MCZ-FALL]; Weber Co. , 12.vi.1937, G. F. Knowlton [1♂, EUB difficilis, BYUC   ]; Unknown Co. , Ut, [1?, litigiosus, MCZ-HORN   ]. VIRGINIA: Alexandria Co. , Alexandria, 1917 [2♂♂ 3♀♀, AMNH]; Arlington Co. , Glencarlyn, 21.vi.1913, Shoemaker [1♀, USNM]; Fairfax Co. , Clifton, 23.vi.1933, J. C. Bridwell [1♂, USNM]; Mt. Vernon , 24.v.1914, A. Wetmore [1♀, USNM]. WASHINGTON: Whitman Co. , Pullman, 23.vi.1947, Esselbaugh [1♀, EUB difficilis, FMNH   ]; same data, except 25.v.1949, N. M. Downie [1♂, EUB difficilis, FMNH   ]. WEST VIRGINIA: Mason Co. , McClintic Wildlife Management, 5.vi.1996, S. M. Clark [1♀, BYUC]; same data, except 2.vi.1998 [2♀♀, BYUC]; same data, except 8.vi.2000, S. M. Clark [3♂♂, RJB carbonarius, BYUC   ]. WYOMING: Campbell Co. , 7 mi N Gillette, Hwy. 14/16, 22.vi.2009, J. Huether & R. Turnbow [1♀, BYUC]; Sheridan Co. , Sheridan LTA, N44°50.011′, W107°01.463, 24-25.vi.2010, B. Kondratieff & B. Heinold [1♂, BYUC]; Weston Co. , 4 mi W Newcastle, 10.vi.1963, D. R. Smith [1♀, NCSU] GoogleMaps   .

EUB

Laboratory of Biology, Faculty of Science, Ehime University

MCZ

Museum of Comparative Zoology

CNC

Canadian National Collection of Insects, Arachnids, and Nematodes

HCF

Universidade Tecnológica Federal do Paraná

OSEC

K.C Emerson Museum

EGR

Eszterházy Károly College

AMNH

American Museum of Natural History

NCSU

North Carolina State University Insect Museum

SMC

Sedgwick Museum

UMRM

W.R. Enns Entomology Museum

OSUC

Oregon State University

INHS

Illinois Natural History Survey

USNM

Smithsonian Institution, National Museum of Natural History

SEMC

University of Kansas - Biodiversity Institute

LSAM

Louisiana State Arthropod Museum

FSCA

Florida State Collection of Arthropods, The Museum of Entomology

CMNH

The Cleveland Museum of Natural History

UMSP

University of Minnesota Insect Collection

UNSM

University of Nebraska State Museum

CUAC

Clemson University Arthropod Collection

SDSU

Severin-McDaniel Insect Collection