STRONGYLOPHTHALMYIIDAE

Papp, L., Merz, B. & Földvári, M., 2006, DIPTERA OF THAILAND A summary of the families and genera with references to the species representations, Acta Zoologica Academiae Scientiarum Hungaricae 52 (2), No. 2, pp. 97-269 : 165-167

STRONGYLOPHTHALMYIIDAE

There are but a few acalyptrate dipterous families, for which we can say that they are rather well known in the Oriental region . One of those exceptions is the family Strongylophthalmyiidae . STEYSKAL ( CDO, 1977) listed 22 Oriental species, although none of them was mentioned from Thailand. SHATALKIN (1996) described eight additional species, two of them from Thailand (both of them found also in our material). FREY’ s (1956) and STEYSKAL’ s (1971) keys, together with SHATALKIN’ s works (1993 etc.) are proper bases for the identification of the Oriental species (only two females in our material ( PF04 /25) was left unnamed) .

We selected and pinned 71 Strongylophthalmyia specimens in 2004. One additional specimen was captured by our colleagues in 2003. The material is rich in species; four of them are new to science.

Strongylophthalmyia brunneipennis (DE MEIJERE, 1914) – 2 f: PF04/42; 1 m: PF04/43. It was described from Java and known also from the Philippines.

Strongylophthalmyia freidbergi SHATALKIN, 1996 – 1 m: PF04/11; 1 m: PF04/14. This is a peculiar species, which was described from NW Chiang Mai (Doi Suket) and from another Thai locality. It is much to our satisfaction, that we managed to capture it again.

Strongylophthalmyia gibbifera SHATALKIN, 1993 – 5 m: PF04/14. It was described from Vietnam.

Strongylophthalmyia lutea (DE MEIJERE, 1914) – 1 m: PF04/43. It was described from Java.

Strongylophthalmyia metatarsata DE MEIJERE, 1919 – 1 m: PF04/42. It was described from Sumatra.

Strongylophthalmyia nigricoxa (DE MEIJERE, 1914) – 1 m: PF04/14; 3 m: PF04/17; 4 m: PF04/19; 1 m: PF04/26; 3 m 2 f: PF04/42; 1 m: Trang Prov., Khao Chong Reserve , rainforest, 22. 11. 2003, river Khao Chong, beating and netting, OROSZ & SZIRÁKI. It was described from Java. It does not seem rare in Thailand. It is a rather characteristic species with its large clypeus, anepisternum shiny black, not setose, male cerci extremely long, slender, longest cercal setae only 2/5 as long as cercus .

Strongylophthalmyia pectinigera SHATALKIN, 1996 – 1 m: PF04/14; 1 m: PF04/17. It was described from NW Chiang Mai (Doi Suket). Now it is reported from other two localities of N Thailand.

Strongylophthalmyia polita (DE MEIJERE, 1914) – 1 f: PF04/14. It was described from Java and known also from Sumatra.

Strongylophthalmyia punctata HENNIG, 1940 ( Figs 60–62 , 132 ) – 4 m: PF04/14; 1 m: PF04/17; 2 m: PF04/19. It was described from Taiwan. The large S-formed process of its 1st flagellomere is very characteristic ( Fig. 132 ). Male palpus extremely broad. Cerci ( Fig. 60 ) more rounded apically than in S. thaii sp. n. Ejaculatory apodeme ( Fig. 61 ) broad fan-shaped. Ventral apical (actually cranial) part of hypandrial complex ( Fig. 62 ) digitiform.

Strongylophthalmyia spinosa FREY, 1956 – 1 m: PF04/14. It was described from Burma. Our specimen does not fit the original description completely. The presence of spiniform black setae on the dorsal side of fore femur is not specific character, since there are a number of species ( palpalis , punctata , thaii ), which possess them. Consequently, that character alone does not define a species.

Strongylophthalmyia crinita group spp. – 1 f: PF04/14; 1 f: PF04/42.

Strongylophthalmyia dorsocentralis L. PAPP , sp. n.

( Figs 129–130 )

Holotype female ( HNHM): THAILAND: Trang Prov., Khao Chong Botanic Garden , along the stream below waterfall, Nov 14, 2004, No. 30, leg. L. PAPP & M. FÖLDVÁRI.

Paratype female (a freshly emerged specimen, legs wrinkled, wing patterrn faint): data same as for holotype.

Measurements in mm: body length 4.67 (holotype), 3.70 (paratype), wing length 3.51, 3.13, wing breadth 1.28, 1.12.

A comparatively large yellow species, with a pair of faint ochre stripes just outside dc lines ( Fig. 130 ).

Head yellow, only ocellar triangle dark. Only 2 pairs of long fronto-orbitals, anterior pair short, lateroclinate, posterior pair long, reclinate. No setae on cheeks and genae but only behind and below eyes. Clypeus extremely high, 0.25 mm, distance of lunule clypeus in sagittal line only 0.31 mm. Ocellar seta long, 0.35 mm. Seta on pedicel only 0.35 mm. Arista bare, 0.70 mm long. Palpus small, almost parallel sided.

Mesonotal setae all brown. 1 weak pprnt, 2 np, 1 presut ia, 1 sa, 1 pa, 1 +3 dc, 1 anepst (rather long), no row of white hairs on centre of anepisternum. Dorsocentrals between the characteristic (countable) ones) are unusually strong ( Fig. 130 ). Only 2, more or less unarranged rows of scattered long acrostichals. No dorsal scutellars, apical scutellars long.

Legs and setae and hairs on legs yellow, dorsal side of fore femur convex, without spines. Fore tibia apically and medially with a comb of adpressed setulae.

Wing ( Fig. 129 ) yellowish with a transverse central and an apical brown spot. Costa and vein R1 yellow, other veins brown (!) . Costa only slightly thickened distally to R1 . Base of R4 +5 thickened and dark brown. Intra-crossvein sections and distal section of M (holotype): 24: 67: 98. Costal sections 78: 101. Calyptrae fumose, fringe rather long, longest 0.34 mm. Haltere ochre, stalk darker, brown .

Female abdomen brown, except for basal 2 tergites, lateral and marginal setae on tergites 0.35 to 0.38 mm long. Female cerci 0.35 mm long, brown, apical hairs only 0.02 mm long.

The specific ephithet refers to the unusually strong dorsocentral setae between the paired

“characteristic” dorsocentrals and the presence of a pair of faint ochre stripes just outside dc lines.

Strongylophthalmyia dorsocentralis sp. n. is an easily recognisable species. It keys to S. lutea DE MEIJERE or to S. immaculata HENNIG both in STEYSKAL’ s (1971) and FREY’ s (1956) keys. Its abdomen is probably yellowish but darkened owing to gut content. Contrarily to S. lutea , it has two large brown spots on wing, its R2+3 is much longer than that of S. lutea .

Figures

Figs 60–65. Strongylophthalmyia spp., male genitalia.60–62 = S. punctata HENNIG: 60 = epandrium and cerci, lateral view, 61 = ejaculatory apodeme, broadest extension, 62 = gonopod, ventral part, broadest view; 63–65 = S. thaii sp. n. paratype: = gonopod, ventral part, broadest view, = epandrium

Figs 129–132. Strongylophthalmia spp. 129–130 = S. dorsocentralis sp. n., holotype female: 129 = wing, 130 = mesonotum dorsally; 131 = S. palpalis sp. n., male palpus, lateral, 132 = S. punctata