Clistoabdominalis nitidifrons ( Becker, 1900 )

FÖLDVÁRI, MIHÁLY, 2013, Taxonomic revision of the Afrotropical species of the tribe Eudorylini (Diptera, Pipunculidae), Zootaxa 3656 (1), pp. 1-121 : 17-18

Clistoabdominalis nitidifrons ( Becker, 1900)

( Figs 5A–E , 40F )

Pipunculus nitidifrons Becker 1900: 241

Pipunculus confusoides Lamb 1922: 412 .—syn. Kehlmaier 2005 a: 23.

Dorylomorpha lini Hardy 1971: 81 .—syn. (with P. confusoides ): De Meyer 1995: 288.

Diagnosis: The basally situated cross-vein, the missing (or at least extremely reduced) pterostigma and the separated eyes in males make this species also very special. GP weakly developed, PG with small protuberances in ventral view.

Type material: Philippines: 1♀ [AT of D. lini ], Philippines : Palawan, 13km N of Puerto Princesa, 18.IV.1968, [hand-written:] “ex grass”, D.E. Hardy Collector; 1♂ [PT of D. lini ], ibid. (both BPBM) . Seychelles: 2♂, 1♀, LT and 2 PLT of P. confusoides , Mahe, '08-9., Seychelles Exp.; Seychelles Is. , Pres by Percy Sladen Trust Cttee., B.M. 1922-157. (all BMNH) . Taiwan: 1♂ [PT of D. lini ], “CHINA: N. TAIWAN, sweeping [on rice], Taipei, 26.VI.1969, [leg.] KSLin”, [separate label:] “# 8a” ( BPBM) .

Male

Head. Third antennal segment acuminate; brownish yellow. Face silvery pollinose. Frons pollinose on lower 1/3, otherwise shining black; eyes do not touch. Frons at least as wide as largest ommatidium at narrowest point.

Occiput, lower half silvery pollinose, upper half brownish.

Thorax. Humeri yellow. Mesonotum (viewed obliquely from front) subshining black; silvery pollinose from the side. Scutellum subshining black, without hairs. Dorsocentral hairs short, pale. Halter yellow.

Legs. Trochanters and femora brown, basal 1/3 of tibiae yellow, otherwise brown. Tarsal segments yellow-brown, last segment brown. Ventroapical row of 5–6 very short, black spines on f2; no spines on other femora. Subapical (distal) spines on first four tibiae present. No anteromedial hairs on 3rd tibia. Hind trochanter without hairs or spines on ventral side. Pulvilli shorter than last tarsal segment.

Wing. Fourth costal section 2 times as long as third costal section. Cross-vein R-M at 1/5–1/6 of discal cell. No pterostigma can be observed on ST (slightly darker in the distal corner of third costal section). One hair on tegula.

Abdomen (already dissected). Viewed obliquely from front first two tergites completely silvery pollinose. Laterally 1–2 minute pale hairs on first tergite. Postabdomen in dorsal view: ST and 2PTs already dissected (T6, S7, EP) visible.

Genitalia. Surstyli small, OS bent outwards; epandrium as long as wide; ST8 broad ( Fig. 5B ). SES uncertain; gonopods weakly developed, small; hypandrium deflected in ventral view ( Fig. 5D ); phallic guide uncertain in lateral view, and with small protuberances in ventral view; phallus very thin; ejaculatory apodeme broad at tip; sperm pump small, globular, phallic sheath present ( Fig. 5D ).

Female

As male except for the following characters. Frons, eyes separated; silver-grey pollinose on lower 1/3, otherwise shining black (the black part is V-shaped). Pulvilli and claws about as long as last tarsal segment. Female ovipositor as in Fig. 40F .

Remarks —Based on the male genitalia this species belongs to Clistoabdominalis , although the pterostigma seems to be missing or at least extremely reduced. One of the male syntypes of P. confusoides is selected here as lectotype to establish an unambiguous species concept.

Distribution— Egypt India, Israel, Japan, Philippines, Reunion Island, Seychelles Islands, Taiwan, Thailand.

Figures

FIGURE 5. Male genitalia of Clistoabdominalis nitidifrons (A–E) and Cli. crassus (F–K).—A: outer surstylus with phallic guide, lateral view, B: dorsal view, C: inner surstylus, lateral view, D: ventral view with phallic guide, sperm pump and ejaculatory apodeme (without ST8), E: surstyli, dorsal view, F: outer surstylus with phallic guide, lateral view, G: dorsal view, H: inner surstylus, lateral view, I: ventral view with phallic guide, (without ST8), J: surstyli, dorsal view, K: sperm pump with ejaculatory apodeme. Scale bars: 0.1 mm

FIGURE 40. Female ovipositors.—A–B: Eudorylas aethiopicus, C–D: Eudorylas bredoi, E: Clistoabdominalis lomholdti, F: Clistoabdominalis nitidifrons, G: Eudorylas falcatus, H–I: Eudorylas flexus, J: Clistoabdominalis crassus, K: Eudorylas setiformis, L: Clistoabdominalis lomholdti, M: Dasydorylas sordidatus, N–O: Eudorylas mikenensis, P–Q: Eudorylas decorus, R–S: Eudorylas galeatus, T: Eudorylas sinuosus, U: Eudorylas wittei, V–W: Dasydorylas evanidus. Scale bar: 0.5 mm