Eudorylas garambensis ( Hardy, 1961 )

FÖLDVÁRI, MIHÁLY, 2013, Taxonomic revision of the Afrotropical species of the tribe Eudorylini (Diptera, Pipunculidae), Zootaxa 3656 (1), pp. 1-121 : 69

Eudorylas garambensis ( Hardy, 1961)

( Figs 22G–L , 39D )

Dorilas (Eudorylas) garambensis Hardy 1961: 138 .

Dorilas (Eudorylas) megacanthus Hardy 1961: 142 — syn. nov.

Diagnosis: Hind trochanter with 2 black spines posterodorsally. Surstyli subsymmetrical, inner tips pointing towards each other; gonopods subsymmetrical with small hairs; phallic guide with two lateral lobes and (in lateral view) a hook; phallus trifid with teeth.

Type material: Democratic Republic of Congo: 1♂, HT, Congo belge P.N.G., Miss[ion] H. De Saeger, II/gd/4, 30-x-1951, Réc. H. De Saeger 2701 ; 1♀, AT, same as HT, II/fd/17, 13-II-1952, No. 3125; 1♀, PT, same as AT, II/ jd/11, 1-IX-1952, No. 4008 ; 1♂ [HT of E. megacanathus ], Congo belge P.N.G., Miss[ion] H. De Saeger, Ndelele/ R, 24-ix-52, H. De Saeger 4075 (all MRAC) .

Other material examined: Botswana: 1♀, Mochudi (bush), 28.05– 9.06.1985, ( MV), leg. and det. De Meyer ; 1♀, same data, 22– 27.05.1985 ; 1♂, Mochudi , 22– 27.05.1985, yellow water trap, leg. De Meyer (all ISNB) . Comoro Islands: 1♀, Moheli Island, 3.10.1983 ; 1♂, same data, 27.09.1983 (both MRAC) . Kenya: 1♂, Ngong Forest Station , IV-13-17-1968, Paul Spangler ( USNM) ; 1♂, same data, 13–18 Jan. 1968, Malaise trap, Krombein & Spangler ( HNHM) .

Male

Head. Third antennal segment acuminate; pale brown. Face silvery pollinose. Frons, upper part black, lower part silvery shining; eyes touching for distance equal to 2.5–3 times ocellar triangle. Occiput, upper half black, lower half silvery grey.

Thorax. Humeri pale brown to dark brown. Mesonotum (viewed obliquely from front) shining black, along anterior margin greyish pollinose; brown from the side with greyish pollinosity. Scutellum shining black, with short, pale hairs. Dorsocentral hairs weakly developed. Halter yellow-brown.

Legs. Trochanters and base of femora yellow, femora yellow (distal 2/3 sometimes brown), knees, tibiae, tarsal segments yellow, last segment brown. Ventroapical row of 7–10 strong, black spines on first four femora; 6–8 spines on 3rd femur. Subapical (distal) spines on first four tibiae present, but weakly developed. One anteromedial hair on 3rd tibia. Hind trochanter with 2 black spines posterodorsally. Pulvilli somewhat shorter than last tarsal segment.

Wing. Fourth costal section 0.7–0.9 times as long as third costal section. Cross-vein R-M at 2/5 of discal cell. Pterostigma fully coloured, proximal border as a line. 1–2 black hairs on tegula.

Abdomen. Viewed obliquely from front tergites shining brown, weakly sclerotised. Hairs very weakly developed. Lateral spines on first tergite: 2–4. Postabdomen in dorsal view: T5 1.1 times as long as ST8. Genitalia without dissection: SS yellow, m.a. elongated.

Genitalia. Surstyli subsymmetrical, inner tips pointing towards each other; epandrium short; ST8 enlarged, broad, m.a. large ( Fig. 22H ). SES borders only clear in lateral view; gonopods subsymmetrical with small hairs ( Fig. 22L ); phallic guide with two lateral lobes and (in lateral view) a hook, and a membranous supporting structure; phallus trifid with teeth; ejaculatory apodeme elongated, broadening in the middle ( Figs 22G, L ); sperm pump flat, simple ( Fig. 22K ).

Female

As male except for the following characters. Frons, eyes separated; silver-grey pollinose on lower half (less so in the median line), shining black on upper half. Pulvilli and claws about 1.5 times as long as last tarsal segment on first leg. One or two erect dark hairs on t3. Tergites on abdomen not weakly sclerotized, two silver patches in the middle of T2 dorsally. Female ovipositor as in Fig. 39D .

Distribution— Botswana, Comoro Islands, Democratic Republic of Congo, Kenya.

Figures

FIGURE 22. Male genitalia of Eudorylas galeatus (A–F) and E. garambensis (G–L).—A: outer surstylus with phallic guide, phallus and outer gonopod, lateral view, B: dorsal view, C: inner surstylus, lateral view, D: sperm pump with ejaculatory apodeme, E: surstyli, dorsal view, F: ventral view with phallic guide, phallus (without ST8), G: outer surstylus with phallic guide, phallus and outer gonopod, lateral view, H: dorsal view, I: inner surstylus, lateral view, J: surstyli, dorsal view, K: sperm pump with ejaculatory apodeme, L: ventral view with phallus and phallic guide (without ST8, epandrium partly). Scale bars: 0.1 mm

FIGURE 39. Female ovipositors.—A: Eudorylas liberia, B: Eudorylas setiformis, C: Eudorylas aemulus, D: Eudorylas garambensis, E: Eudorylas remiformis, F: Eudorylas mutillatus (E. eremnoptera), G–H: Eudorylas definitus (E. excisus), I: Eudorylas facetus, J–K: Dasydorylas quasidorsalis, L: Eudorylas amuscarinum, M: Eudorylas abdominalis, N–O: Clistoabdominalis namibiensis sp. nov., P: Eudorylas mutillatus, Q: Eudorylas femoralis sp. nov., R: Eudorylas ghesquierei, S: Dasydorylas okongoensis sp. nov., T–U: Eudorylas angustus, V: Eudorylas excisus. Scale bar: 0.5 mm