Achmonia simulator, Janák, Jiří & Bordoni, Arnaldo, 2014

Janák, Jiří & Bordoni, Arnaldo, 2014, Revision of the genus Achmonia of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae), Zootaxa 3872 (3), pp. 257-274 : 267-269

Achmonia simulator sp. nov.

( Figs. 4, 7, 9 , 19 , 37 , 51–63 )

Type locality. D. R. Congo, Uele, Itimbiri, Dingila.

Type material. Holotype ♂: DEMOCRATIC REPUBLIC OF CONGO “Musée du Congo, Uele - Itimbiri Dingila, 6-X-1932, J. Vrydagh” ( MRAC). Paratypes: CAMEROUN: 1 ♂: “ Kamerun, Joko, L. Colin, VIII, 11”, ( JJRC); 1 ♀: “Joko, Kamerun, Heyne, Berlin W” ( MFNB); 1 ♀: “ Kamerun, Joko” ( MFNB); 1 ♀: “ Cameroon, Jacum, L. Colin VI.1911 ” ( MFNB); CENTRAL AFRICAN REPUBLIC: 1 ♂: “Uamgebiet, 1-10.IV.1914, Bosum [= Bozoum], Tessmann S.” ( ABFI); GUINEA BISSAU: 1 ♂: “ Guinea-Bissau, Bolama, L. Fea VI- XII.1899 ” ( ABFI); NIGERIA: 1 ♂, 2 ♀: “ Nigeria, Umg Jos/Vos, V. Assing 5.XI–2.XII.1984 ” ( VAHG, ABFI); ZAMBIA: 1 ♀: “ Zambia NC, Mkushi env. E, 16– 18.12.2004, Snížek + Tichý lgt.” ( JJRC); 1 ♀: “ Zambia NW, 27 km N Kasempa, 10.12.2004, Snížek & Tichý” ( JJRC), 1 ♂: “ Zambia, Central. prov., 30 km S Kapiri Mposhi, 6.XII.2009, M. Bednařík leg., 1150m ” ( JJRC).

Description. Body length ( Fig. 4 ) 10.5–14.0 mm; length from anterior margin of head to posterior margin of elytra: 5.5–7.5 mm. Black, elytra brownish with yellowish spots near humeral angles and posterior margin; abdomen brownish black with genital segment yellow; antennae and legs brown. Head and pronotum with micropunctation. Head with postocular punctures. Labrum quadrilobate, with moderately prominent inner lobes ( Fig. 19 ). Head slightly to distinctly widened posteriad, punctation as in Figs. 9 , 37 . Elytra as long as pronotum, posteriad, dilated and wider than pronotum; surface with fine punctation, arranged in three series. Abdomen with transverse micro-striation and fine, not sparse punctation on the sides.

Male. Tergite and sternite of the male genital segment as in Figs. 52, 53 , 62 . Aedeagus ( Figs. 54, 55, 58, 61 ) length 1.05–1.15 mm, with median lobe long and narrow, pointed ( Figs. 57, 60 ); paramere long and slender ( Figs. 56, 59 ); internal sac very narrow, covered by fine and sparse scales.

Ratios (n = 9): HL/HW: 0.94–1.03 (M 0.99); TeL/EyL: 1.85–2.26 (M 2.03); PL/PW: 1.19–1.24 (1.21); EL/EW: 1.01–1.08 (M 1.05); PW/HW: 0.83–0.95 (M 0.89); EW/PW: 1.09–1.19 (M 1.11); EL/PL: 0.93–0.99 (M 0.96); A5L/A5W: 0.59–0.68 (M 0.63); A10L/A10W: 0.54–0.64 (M 0.60).

Differential diagnosis. Achmonia simulator sp. nov. differs from other Afrotropical species of the genus by the surface among the setiferous postocular punctures, which is without punctures and by the narrow and pointed median lobe of the aedeagus.

Etymology. The name of the species is a noun in apposition—Latin word “ simulator ” due to its similarity with A. flavomarginata .

Distribution. The new species is recorded from D. R. Congo, Cameroon, Central African Republic, Guinea Bissau, Nigeria and Zambia ( Fig. 1).

Remarks. The differences in the shape of the paramere in Fig. 61 compared to the typical form shown in Figs. 56 and 59 are partly due to a more schematic style of drawing (Arnaldo Bordoni) and partly due to intraspecific variability.

Figures

FIGURES 3 – 12. 3, 8, 12, Achmonia carinata sp. nov., HT male; 4, 7, 9, A. simulator sp. nov., PT male, 4, 9 from Mposhi, 7 from Joko; 5, 10, A. amabilis (Boheman), female from Ithala; 6, 11, A. congoensis (Bernh.), male from P. N. Garamba. 3 – 6, habitus; 7, lateral part of pronotum, ventral; 8 – 11, head; 12, pronotum and elytra. SL, superior line; IL, inferior line; EL, elytral length; p, postocular puncture. Scales 1 mm (3 – 6) and 0.5 mm (7 – 12).

FIGURES 13 – 24. 13, 14, 21, 22, Achmonia doloc (Bord.), HT from Java; 15, A. eppelsheimi (Bernh. & Schub.) from Nepal; 16, A. shibatai (Bord.), HT from Taiwan; 17, A. nigra (Bord.), HT from China; 18, 23, 24, A. amabilis (Boh.) from South Africa; 19, A. simulator sp. nov., PT from Mposhi; 20, A. carinata sp. nov., HT. 13, head and pronotum; 14 – 20, labrum; 21, 23, maxillary palpus; 22, 24, labial palpus. 13, 14, 21, 22 from Bordoni (2004 a), 15 from Bordoni (2004 b), 16 from Bordoni (2010), 17 from Bordoni (2009). Scale 0.1 mm; 13, 15 and 17 without scale.

FIGURES 25 – 37. 25 – 31, Achmonia knirschi sp. nov., HT male; 32 – 34, A. carinata sp. nov., 32, PT female, 33, 34, HT male; 35, A. flavomarginata (Bernh.), HT male; 36, A. kapangana sp. nov., HT male; 37, A. simulator sp. nov., HT male. 25, 32, 35 – 37, head; 26, pronotum dorsal; 27, aedeagus; 28, median lobe of aedeagus; 29, paramere; 30, 33, male tergite 10; 31, 34, male sternite 9. Scales 1 mm (25 = 26; 32 = 35 – 37), 0.5 mm (27 = 30, 31, 33, 34) and 0.1 mm (28 = 29).

FIGURES 38 – 53. 38 – 41, Achmonia carinata sp. nov., HT male; 42 – 44, A. kapangana sp. nov., HT male; 45 – 50, A. flavomarginata (Bernh.), LT male; 51 – 53, A. simulator sp. nov., HT male. 38, 45, aedeagus lateral; 39, 42, 46, aedeagus dorsal; 40, 43, 47, paramere; 41, 44, 48, median lobe of aedeagus; 49, 52, male sternite 9; 50, 53, male tergite 10; 51, pronotum dorsal. B, basal bulbus; ML, median lobe. Scales 0.5 mm (38 = 39; 42 = 45, 46, 49, 50, 52, 53; 51) and 0.1 mm (40 = 41; 47 = 43, 44, 48).

FIGURES 54 – 68. 54 – 63, Achmonia simulator sp. nov., 54 – 57, 63, PT male from Joko, 58 – 61, HT male, 62, PT from Bolama; 64 – 67, A. amabilis (Boheman), 64, 66, 67, LT male, 65, female from Ithala; 68, A. congoensis (Bernh.), male from Malo. 54, aedeagus lateral; 55, 58, 61, aedeagus dorsal; 56, 59, paramere; 57, 60, median lobe of aedeagus; 62, sternite 9 and sclerites of tergite 10; 63, left metatibia and metatarsus; 64, 68, head; 66, sternite 9; 67, tergite 10; 65, lateral part of head. Scales 1 mm (64 = 65, 68), 0.5 mm (54 = 55, 58, 61, 62; 63; 66 = 67) and 0.1 mm (56 = 57, 59, 60).