Tomaspisinella Lallemand

Andrew Hamilton, K. G., 2016, Neotropical spittlebugs related to Neaenini (Hemiptera, Cercopidae) and the origins of subfamily Cercopinae, Zootaxa 4169 (2), pp. 201-250: 223-224

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Tomaspisinella Lallemand


Tomaspisinella Lallemand   , redefined

Tomaspisinella Lallemand, 1927: 117   . Type-species by original designation: T. parva Lallemand, 1927   .

Hemitomaspis Lallemand, 1949: 31   , syn.nov. Type-species by original designation: Tomaspis caligata Jacobi, 1908   .

Distribution. Neotropical, from Costa Rica south to Venezuela.

Diagnosis. Superficially similar to Menytes   ( Fig. 14 View FIGURES 14 – 15 A) but without sulcate face; color usually black to brown (except jocosa   sp. nov.), body shorter, broader across the head, with convex tegmina ( Fig. 35 View FIGURES 25 – 35 ) and distinctive antenna ( Figs 19 View FIGURES 17 – 19 A –C) with postpedicel retracted into pedicel, as in Orthoraphini (Aphrophorinae) and basiconic sensillum elongate and setiform, as in Menytes   and Simorhina (Neaenini)   and Sphenorhina Amyot & Serville (Ischnorhinini)   .

Description. Head distinctly to slightly narrower than pronotum ( Carvalho & Webb 2005, figs 280–281); eyes varying from globose to transverse; crown short, weakly concave; antennal ledge arched above coronal margin; frons convexly inflated. Pronotum with anterior margin steeply declivous ( Fig. 5 View FIGURES 1 – 8 C), anterior border weakly to distinctly bowed; lateral margins as long as eyes to much shorter than half length of eyes, surface variable from medially carinate and transversely rugulose ( Fig. 26 View FIGURES 25 – 35 ) to smooth with scattered pits ( Fig. 27 View FIGURES 25 – 35 ) or densely setose ( Fig. 35 View FIGURES 25 – 35 , Merinx subg. nov.), the nominate subgenus having fine setae ( Fig. 5 View FIGURES 1 – 8 C). Tegmina convex, shiny to hirsute; hind wing with 3 apical cells (Cu is unbranched; appendix large, even width around tip of wing. Fore femora distinctly longer than hind femora. Hind tibiae with 2 spines on outer edge, both long in smallest species, but basal one tiny in largest species; hind tibial pectin with 8–9 spines; hind basitarsal pecten of 6–7 black-tipped spines, that of second tarsomere with 8–9 such spines (hind legs missing from type of T. parva   ); arolia variable from much shorter than claws (in the type-species) as in Ischnorhinini, to longer than the claws (in Merinx subg. nov.). Male pygofer usually elongate and deeply notched dorsally on each side of anal tube ( Figs 28–35 View FIGURES 25 – 35 A) as in Menytes   ; subgenital plates absent or broad, fused to pygofer, nearly vertical, upper angles with decurved, pointed tips, sometimes with long, median processes ( Fig. 34 View FIGURES 25 – 35 D); styles unarmed or absent; phallobase short, sometimes fused to theca to form aedeagus; shaft unarmed or with paired, retrorse processes ( Figs 28 View FIGURES 25 – 35 C, 34–35C).

Included species. The original description included Tomaspisinella caligata (Jacobi)   T. parva Lallemand   , T. minuscula (Jacobi)   and 2 species now assigned to Zuata   . Carvalho & Webb (2005) added T. apicifasciata (Fowler)   , known only from the female type, and these identities (except that of minuscula   ) have been confirmed by examination of the antennae. T. punctatissima   (Stål: Lepyronia   ) comb.nov. and T. ignobilis (Fowler)   , comb.nov. were formerly placed in Hemitomaspis Lallemand   and are here transferred to Tomaspisinella   . These together with 6 new species increase the genus to 12 species. The typical subgenus appears to be paraphyletic with respect to 3 other apomorphic subgenera, with T. lucifer   sp. nov. intermediate in pronotal characters and T. diabolos   sp. nov. intermediate in genital characters.












Tomaspisinella Lallemand

Andrew Hamilton, K. G. 2016


Lallemand 1949: 31


Lallemand 1927: 117