Microclimax, Andrew Hamilton, K. G., 2016

Andrew Hamilton, K. G., 2016, Neotropical spittlebugs related to Neaenini (Hemiptera, Cercopidae) and the origins of subfamily Cercopinae, Zootaxa 4169 (2), pp. 201-250: 210

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gen. nov.

Microclimax   gen. nov.

Type-species: Tomaspis luteosignatus Valdés Ragués (1910) sensu Metcalf & Bruner (1944)   , the only included species.

Etymology. micro- small, klimax (f), ladder, referring to the veins of the costal margin.

Distribution. Cuba.

Diagnosis. Tegmina each with 2 large anteapical cells, without small, round central anteapical cell, costal margin with ladder of 5–6 perpendicular crossveins, apical cells large, separated by unbranched veins.

Description. Head distinctly narrower than pronotum; crown weakly impressed transversely, with tip upturned, bearing grooves from angles of tylus to ocelli; anterior margin of crown not carinate; antenna with postpedicel bearing numerous large pits and a beaklike basiconic sensillum ( Fig. 24 View FIGURES 24 F); eye round, as wide as lateral margins of pronotum. Tegmen with 5 perpendicular crossveins connecting outer anteapical cell to costa. Costal margin of hind wing with triangular lobe as in Liparonotum   , but its 3 stout hooks are recumbent and equally spaced. Hind tibiae each with double pecten of 9 black-tipped spines, hind tarsal pectens with 9 black-tipped spines on basomere, 11 close-set spines in 2nd tarsomere. Male pygofer dorsoventrally compressed and armed with a spine on posterior margin as in Microsargane   ( Fig. 50 View FIGURES 46 – 54 A–B), but longer, extending nearly to tips of subgenital plates ( Fig. 60 View FIGURES 55 – 60 A); anal tube weakly sclerotized, surrounded by rounded lobes on the dorsal margin of the pygofer; subgenital plate elongate, tapered to blunt tip ( Fig. 60 View FIGURES 55 – 60 D); phallobase short ( Fig. 60 View FIGURES 55 – 60 B) and very broad, almost as wide as subgenital plates and tapered caudad; style broad, apical half bearing very long setae and deeply notched just before robust apical spine directed dorsad ( Fig. 60 View FIGURES 55 – 60 A), in dorsal aspect with 2 short teeth on upper edge ( Fig. 60 View FIGURES 55 – 60 G); theca tubular, weakly laterally compressed, bearing some 90 spinelike setae in tufts flanking gonopore ( Fig. 60 View FIGURES 55 – 60 B–C), the latter slitlike, extending half length of shaft on anterior surface. Ovipositor base preceded by broad, rectangular plate arched over 1st valvifers, itself preceded by bulbous, strongly sclerotized body near sternum, open on posterior face and along midline of upper side.

Remarks. Similar to Liparonotum   in the length of the lateral margins of the pronotum, the shape of the costal lobe on the hind wing and the spiny theca, but the antennal basiconic sensillum is broad rather than spinelike, the lora are narrower, the tegminal venation is quite different and the male pygofer bears a very prominent lateral spine.

Only one species is known.