Hydrochus, Leach, 1817

Perkins, Philip D., 2019, Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae), Zootaxa 4708 (1), pp. 1-59 : 8-10

Hydrochus VAIS collecting events, 2006-2012:

AS-06-009: Venezuela: Apure State: Road between San Fernando and Rio Capanaparo, 0.5 km N. Rio Claro, 67° 38.69’ W, 7° 10.162’ N, 50m, 01/04/2006, leg. Short & Torres, roadside ditch/swale

AS-06-013: Venezuela: Amazonas State: Road between Puerto Ayacucho and Samariapo, 67° 45.358’ W, 5° 20.497’ N, 72m, 01/06/2006, leg. Short, small stream in road culvert

AS-06-019: Venezuela: Apure State: Road between Rio Orinoco and Rio Cinaruco, 67° 32.604’ W, 6° 30.9’ N, 68m, 01/08/2006, leg. Short & Torres, Morichal

AS-08-081a: Venezuela: Amazonas State: Puerto Ayacucho, approx 15 km S, nr. Campamento Canturama, 67° 36.109’ W, 5° 30.623’ N, 100-110m, 08/08/2008, leg. Short, pool at base of large rock outcrop

VZ08- 1230-02X: Venezuela: Zulia State: Quebrada Riencito, nr, 72.30837° W, 10.85498° N, 81m, 12/30/2008, leg. Short, Camacho, García, Marshy pond along road

VZ09-0104-01A: Venezuela: Aragua State: Henri Pittier National Park, Rio La Trilla, 67.74250° W, 10.37319° N, 295m, 01/04/2009, leg. Short, García, Camacho, Miller

VZ09-0108- 03X: Venezuela: Guarico State: N of Palenque, 66° 59.595’ W, 9° 6.794’ N, 152m, 01/08/2009, leg. Short, Camacho, García, Joly, Miller, stream margins

VZ09-0109- 01X: Venezuela: Guarico State: Las Mercedes, approx 65 km S, 66° 22.602’ W, 8° 31.705’ N, 145m, 01/09/2009, leg. Short, García, Camacho, Miller, large vegetated lagoons

VZ09-0110-01A: Venezuela: Guarico State: San Nicolasito Field Station, Rio Aguaro, 66° 26.228’ W, 8° 6.226’ N, 52m, 01/10/2009, leg. Short, Camacho, García, Joly, Miller, along river

VZ09-0110- 02X: Venezuela: Guarico State: San Nicolasito Field Station, morichal in front of station, 66° 24.459’ W, 8° 8.296’ N, 50m, 01/10/2009, leg. Short & Miller, morichal margin and backwaters

VZ09-0111-01G2: Venezuela: Guarico State: San Nicolasito, marsh E of, 66° 20.598’ W, 8° 8.713’ N, 74m, 01/11/2009, leg. García

VZ09-0111-01M3: Venezuela: Guarico State: San Nicolasito, marsh E of, 66° 20.598’ W, 8° 8.713’ N, 74m, 01/11/2009, leg. Miller

VZ09-0111- 02X: Venezuela: Bolivar State: between Caicara & Los Pijiguaos, 66° 17.904’ W, 7° 20.992’ N, 62m, 01/11/2009, leg. Short, García, Miller, various lentic habitats: pond and lagoon

VZ09-0112-01A: Venezuela: Bolivar State: Los Pijiguaos, outcrop/morichal, 66° 49.238’ W, 6° 35.617’ N, 80m, 01/12/2009, leg. Short, Camacho, García, Joly, Miller, morichal margin

VZ09-0112-01B: Venezuela: Bolivar State: Los Pijiguaos, outcrop/morichal, 66° 49.238’ W, 6° 35.617’ N, 60- 80m, 01/12/2009, leg. Short, Camacho, García, Joly, Miller, algae on rocky margin of morichal

VZ09-0112-01C: Venezuela: Bolivar State: Los Pijiguaos, outcrop/morichal, 66° 49.238’ W, 6° 35.617’ N, 60- 80m, 01/12/2009, leg. Short & Miller, detrital pools

VZ09-0112-02A: Venezuela: Bolivar State: Rio Caripito, 67.02912° W, 6.58694° N, m, 01/12/2009, leg. Short & Miller, river margin

VZ09-0112-02B: Venezuela: Bolivar State: Rio Caripito, 67.02912° W, 6.58694° N, m, 01/12/2009, leg. Short & Miller, detrital pool

VZ09-0113- 01X: Venezuela: Bolivar State: El Burro, approx 25 km E, 67° 14.467’ W, 6° 13.059’ N, 62m, 01/13/2009, leg. Short, Camacho, García, Joly, Miller, rocky morichal

VZ09-0115-03A: Venezuela: Amazonas State: Communidad Porvenir, just S of, 67° 45.315’ W, 5° 20.514’ N, 87m, 01/15/2009, leg. Short & García, pool in culvert

VZ09-0115-03B: Venezuela: Amazonas State: Communidad Porvenir, just S of, 67° 45.315’ W, 5° 20.514’ N, 87m, 01/15/2009, leg. Short & Miller, small streamlet

VZ09-0117- 01X: Venezuela: Apure State: Road between Rio Orinoco and Rio Cinaruco, 67° 32.604’ W, 6° 30.9’ N, 68m, 01/17/2009, leg. Short, Camacho, Miller, Morichal margin

VZ09-0117- 03X: Venezuela: Apure State: Road between Rio Capanaparo and Rio Cinaruco, Medanos de la Soledad, 67° 43.868’ W, 7° 20.175’ N, 49m, 01/17/2009, leg. Short, Camacho, Miller, marshy area

VZ09-0118- 02X: Venezuela: Apure State: Mantecal, approx 10 km W, side road, 69° 3.679’ W, 7° 37.298’ N, 83m, 01/18/2009, leg. Short, Camacho, Miller, marsh and culvert pool

VZ09-0118- 03X: Venezuela: Apure State: between La Ye and Bruzual, 69° 18.004’ W, 7° 38.660’ N, 90m, 01/18/2009, leg. Short, Camacho, Miller, roadside lagoon

VZ09-0118- 04X: Venezuela: Apure State: Bruzual, edge of town, 69° 20.530’ W, 8° 2.534’ N, 83m, 01/18/2009, leg. Short, Camacho, Miller, large marsh

VZ09-0119- 01X: Venezuela: Barinas State: Libertad, E of, along side gravel road, 69° 35.202’ W, 8° 25.773’ N, 106m, 01/19/2009, leg. Short, Camacho, Miller, forested canal

VZ09-0119-02B: Venezuela: Portuguesa State: Guanare, N of, Rio Guanare, 69° 48.961’ W, 9° 2.508’ N, 185m, 01/19/2009, leg. García

VZ09-0119-02C: Venezuela: Portuguesa State: Guanare, N of Rio Guanare, 69° 48.961’ W, 9° 2.508’ N, 185m, 01/19/2009, leg. Short, side pools in rocky flood plain

VZ09-0710-03B: Venezuela: Falcon State: Tocopero, SE of, 69° 13.109’ W, 11° 26.922’ N, 12m, 07/10/2009, leg. Short, Camacho, Inciarte, García, Gustafson, Shepard, Sites, Large, partially shaded pond with lots of emergent vegetation

VZ09-0714-05A: Venezuela: Barinas State: Los Pasitos, E, 70° 32.194’ W, 8° 28.477’ N, 205m, 07/14/2009, leg. Short, Camacho, Inciarte, García, Gustafson, Shepard, Sites, marshy pond with lots of vegetation on large hacienda

VZ09-0716-05A: Venezuela: Tachira State: Chorro el Indio, 72° 12.725’ W, 7° 42.986’ N, 1389m, 07/16/2009, leg. Short, Camacho, Inciarte, Gustafson, Shepard, Sites, margins of small pond

VZ09-0716-07A: Venezuela: Tachira State: El Tama National Park, by INPARQUES office, 72° 10.340’ W, 7° 35.038’ N, 472m, 07/16/2009, leg. Short, Camacho, Inciarte, Gustafson, Sites, light, mercury vapor

VZ09-0812-02A: Venezuela: Anzoategui State: Transect 1, 64° 8.45’ W, 9° 21.03’ N, 327m, 08/12/2009, leg. Cordero, permanent pond with grass

VZ10-0129-04A: Venezuela: Sucre State: El Pilar, approx 5 km SE, 63° 7.070’ W, 10° 31.419’ N, 2m, 01/29/2010, leg. Short & García, marsh/swamp along road

VZ10-0130-02A: Venezuela: Sucre State: Irapa, nr, river crossing at hwy 9, 62° 35.149’ W, 10° 35.532’ N, 14m, 01/30/2010, leg. Short, García, Joly, mud puddles near river

VZ10-0131-03B: Venezuela: Monagas State: Rio Azagua, 63° 8.548’ W, 10° 0.840’ N, 38m, 01/31/2010, leg. Short & García, side pools

VZ10-0202-01B: Venezuela: Monagas State: Maturin, S of, river crossing, 63° 8.295’ W, 9° 36.591’ N, 45m, 02/02/2010, leg. Short & García, vegetated river/morichal margin

VZ10-0202-03A: Venezuela: Monagas State: Morichal Largo & Temblador, small pond between, 62° 43.618’ W, 9° 5.798’ N, 29m, 02/02/2010, leg. Short, García, Joly, margins of vegetated pond

VZ10-0203-01A: Venezuela: Delta Amacuro State: Between Tucupita & Los Guires, 61° 54.610’ W, 9° 10.504’ N, 8m, 02/03/2010, leg. Short & García, marsh by road

VZ10-0203-02A: Venezuela: Delta Amacuro State: between Tucupita & Temblador, small pond along road, 62° 14.306’ W, 8° 46.439’ N, 19m, 02/03/2010, leg. Short, García, Joly, margins of vegetated pond

VZ10-0203-03A: Venezuela: Anzoategui State: El Tigre, N of, river along highway, 64° 19.445’ W, 9° 5.808’ N, 236m, 02/03/2010, leg. García, shaded side margins without vegetation

VZ10-0204-01A: Venezuela: Anzoategui State: Zaraza, E of, ranch lagoon, 65° 11.769’ W, 9° 21.186’ N, 68m, 02/04/2010, leg. Short & García, vegetated lagoon margin

VZ10-0705-02B: Venezuela: Guarico State: San Nicolasito Field Station, morichal in front of station, 66° 24.459’ W, 8° 8.296’ N, 50m, 07/05/2010, leg. Short, Arias, Tellez, HG vapor and UV lights

VZ10-0706-01A: Venezuela: Guarico State: Santa Rita, 2.6 km W, 66° 16.685’ W, 8° 7.261’ N, 73m, 07/06/2010, leg. Short & Tellez, flooded area by road

VZ10-0717-02A: Venezuela: Bolivar State: Gran Sabana, N Santa Elena, Rio Guara at Rt 10, 61° 5.679’ W, 4° 37.362’ N, 876m, 07/17/2010, leg. Short, Arias, Tellez, marshy area in river floodplain

VZ10-0719-01A: Venezuela: Monagas State: Chaguaramas, 4 km S, 62° 45.918’ W, 8° 38.079’ N, 124m, 07/19/2010, leg. Short, Arias, Tellez, roadside muddy puddle

VZ12-0120-03A: Venezuela: Cojedes State: El Pao, approx 7.5 km, Rio Caiman Grande at San Brano, 68° 11.860’ W, 9° 39.246’ N, 137m, 01/20/2012, leg. Short, Arias, Gustafson, stream margins

VZ12-0121-02A: Venezuela: Cojedes State: La Galera, approx 2 km S, between El Pao & El Baul, 68° 9.908’ W, 9° 32.930’ N, 126m, 01/21/2012, leg. Short, Arias, Gustafson, marsh/pond

VZ12-0121-03A: Venezuela: Cojedes State: El Baul, 5 km S, large marsh, 68° 19.304’ W, 8° 54.008’ N, 70m, 01/21/2012, leg. Short, Arias, Gustafson, marsh

VZ12-0122-01A: Venezuela: Cojedes State: Aparicion, at highway, lagoon/pond, 69° 23.062’ W, 9° 22.268’ N, 213m, 01/22/2012, leg. Short, Arias, Gustafson, pond

VZ12-0125-01A: Venezuela: Barinas State: Ciudad Bolivia, approx 20 km S, small stream, 70° 23.867’ W, 8° 16.954’ N, 149m, 01/25/2012, leg. Short, Arias, Gustafson, stream margins

VZ12-0125-02A: Venezuela: Barinas State: Ciudad Bolivia, approx 13 km SE, large Hacienda, 70° 28.238’ W, 8° 19.394’ N, 173m, 01/25/2012, leg. Short, Arias, Gustafson, marsh

VZ12-0126-01B: Venezuela: Barinas State: Santa Barbara, E of, Rio Santa Barbara, 71° 11.188’ W, 7° 50.028’ N, 177m, 01/26/2012, leg. Short, Arias, Gustafson, big sidepool of river

VZ12-0126-02A: Venezuela: Tachira State: La Pedrera, 10 km E, Mata de Limon, small lagoon on finca, 71° 29.322’ W, 7° 30.172’ N, 185m, 01/26/2012, leg. Short, Arias, Gustafson, marsh

VZ12-0127-01A: Venezuela: Zulia State: Puente del Zulia, lagoon on finca, 72° 20.163’ W, 8° 33.108’ N, 47m, 01/27/2012, leg. Short, Arias, Gustafson, large lagoon

VZ12-0127-02A: Venezuela: Zulia State: Encontrados, approx 3 km SE, 72° 12.734’ W, 9° 1.982’ N, 19m, 01/27/2012, leg. Short, Arias, Gustafson, small lagoon

VZ12-0128-06A: Venezuela: Trujillo State: La Cieba, approx 10 km E, by cemetery, 70° 57.345’ W, 9° 28.505’ N, 18m, 01/28/2012, leg. Short, Arias, Gustafson, lagoon

Following are the complete names of the collectors in the VAIS project: Quintin Arias, Jesus Camacho, Ruben Cordero, Mauricio García, Grey Gustafson, L. J. Joly, Kelly B. Miller, Andrew E. Z. Short, William D. Shepard, Robert Sites, Monica Tellez, and Phil Torres.

Locality data for specimens from other collections are given verbatim in the species accounts, sometimes with very slight changes, such as punctuation, for clarity.

Some duplicate specimens from long series will be deposited in the MCZ.