Clydonium porteri Khalaim, 2023

Khalaim, Andrey I. & Ruíz-Cancino, Enrique, 2023, Darwin wasps of the subfamily Pimplinae (Hymenoptera: Ichneumonidae) of Mexico: Camptotypus genus-group, Zootaxa 5330 (1), pp. 47-72 : 50-51

3. Clydonium porteri Khalaim , sp. nov.

( Fig. 7 )

Material examined. Holotype female ( FSCA), Mexico, San Luis Potosí, 12 km S of Valles, Ruta 85, Hotel Covadonga , 27–29.vi.1981, coll. C. Porter & L.A. Stange.

Paratypes. MEXICO. Tamaulipas: 1 ♀ ( FSCA) 80 km S of Cd. Victoria, La Gloria, Ruta 85, 13.viii.1984, coll. L. Stange & C. Porter. Chiapas: 1 ♀ ( EMEC) 6 mi S of Puebla Nueva , 20.iii.1953, coll. S.C. Bechtel & E.I. Schlinger. GUATEMALA. 1 ♀ ( AEIC) Amatitlán , 7.ii.1905, coll. Jas. S. Nine.

Description. Female. Mandible stout, with upper tooth slightly shorter than lower tooth. Malar space about 0.3× as long as basal mandibular width. Face smooth and shining, finely and sparsely punctate, 1.25× as wide as long. Head in dorsal view with genae strongly and evenly rounded behind eyes. Posterior ocellus separated from eye by 1.15× its own maximum diameter.

Pronotum in profile long, polished, mediodorsally with anterior margin reflexed backwards into a sharp point. Epomia entirely absent. Mesoscutum polished, mostly glabrous, with fine setae in anterior part. Notaulus very weakly impressed. Mesopleuron polished, impunctate in upper part and finely punctate in lower part. Epicnemial carina present ventrally, not reaching up to level of lower corner of pronotum. Epicnemium without secondary carina. Metapleuron almost flat, polished, with very fine punctures in posterior 0.3. Submetapleural carina absent, but with weak ridge anteriorly. Propodeum in profile evenly rounded, smooth, with fine punctures laterally. Pleural carina discernible anteriorly as groove and absent posteriorly (behind propodeal spiracle).

Fore wing length 9.0 mm. Hind wing with first abscissa of nervellus straight, vertical, 1.8× as long as second abscissa.

Metasoma stout. First tergite almost as long as posteriorly broad, with smooth central area that is defined laterally by strong subparallel lateromedian carinae which extend back almost to hind margin of tergite; dorsolateral carinae present near hind margin of first tergite, not extending forwards to spiracles. Second tergite with median indentation on anterior margin strongly impressed. Second tergite smooth, with scattered punctures centrally and laterally, with strong oblique lateral grooves. Tergites 3–5 similar but with lateral grooves more transverse than longitudinal. Ovipositor projecting beyond apex of metasoma by 1.6× length of hind tibia. Ovipositor straight, very strongly compressed laterally, apex of lower valve with scabrous area just proximal to teeth.

Head and antenna black; the apical flagellomere brown. Mesosoma black with mesoscutum reddish brown (sometimes extreme anterior end of mesoscutum fuscous). Legs entirely black. Metasoma reddish brown with extreme apex (tergite 7) black; ovipositor sheath black, apically brownish. Wings strongly darkened with black, pterostigma black.

Male. Unknown.

Variation. Some paratypes are paler, with mesosoma dark reddish brown and propodeum dorsally with vague reddish marks; one female also with pronotum with a broad reddish stripe on upper margin in its posterior half. However, all these specimens have mesoscutum red or reddish brown, distinctly paler and strongly contrasting with remaining black mesosoma.

Etymology. This species is named in honour of Charles C. Porter, in recognition of his outstanding contribution to the taxonomy of Neotropical Ichneumonidae .

Distribution. Mexico (Tamaulipas, San Luis Potosí, Chiapas), Guatemala.

Comparison. The new species is very similar to C. quintanillai but differs by the reddish brown mesoscutum ( Fig. 7 ) (black in C. quintanillai ). This feature is found to work well for all examined specimens of C. porteri sp. nov. and C. quintanillai , with no intermediate forms.

Figures

FIGURES 6–9. Clydonium quintanillai, ♀ (Tamaulipas): 6—metasomal tergites 2–6, dorsal. Clydonium porteri sp. nov., holotype, ♀: 7—mesosoma, lateral. Odontopimpla fasciata, ♀ (from Gauld 1991): 8—hind femur, lateral; 9—apex of ovipositor, lateral.