Cinetomorpha longisetosa Ott & Harvey, 2019

Ott, Ricardo, Ubick, Darrell, Bonaldo, Alexandre B., Brescovit, Antonio D. & Harvey, Mark S., 2019, A revision of the new world goblin spider genus Cinetomorpha Simon, 1892 revalidated from Gamasomorpha Karsch, 1881 (Araneae, Oonopidae, Oonopinae), Zootaxa 4641 (1), pp. 1-152 : 69

Cinetomorpha longisetosa Ott & Harvey , sp. nov.

Figures 49A ̅51F

Type material. Holotype male from La Selva , 10.43333°, -84.01666°, Heredia, Costa Rica, May 11, 2000, C. Viquez, deposited in INBIO PBI_OON 31121 . Paratype: La Selva, 150m, 10.43333°, -84.01666°, Heredia, Costa Rica, Sept. 1, 1992, C. Viquez, 1♀ ( INBIO PBI _ OON 31123) .

Etymology. The specific name is an adjective referring to the long and strong setae in the inner paturon of male chelicerae.

Diagnosis. This species resembles C. puberula comb. rest. and C. zero sp. nov. bythe shape of the suprapedicelar region with two strong downwards directed teeth ( Figs 32B , 34F, 49G, 51E, 60G, 63H), can be distinguished from the first by the punctuated pattern of the sternum without any groove ( Figs 31F , 33A , 49F ) and from the last species by the large ridge connecting the two strong suprapedicelar teeth ( Figs 49G , 51E , 60G , 63H ). Males of this species are also close to C. loreto sp. nov. by the the presence of long and strong sclerotized setae in the inner paturon of male chelicerae ( Figs 49E ̅F), character not possessed by any other species of the group, however can be distinguished from the last species by the shape of the suprapedicelar ridge, which in C. loreto sp. nov. does not present downwards directed teeth ( Figs 52G , 54G ). Female internal genitalia resembles also C. baja sp. nov. ( Fig. 42H ), with domed anterior area and squared posterior area, can be distinguished from the last by the trapezoidal chitinized wall area, by the forward positioned receptaculum and the very right-angled posterior corners of the internal genitalia ( Fig. 51F ).

Description. Male ( PBI _OON 31121). Total length: 1.73. CEPHALOTHORAX: Carapace orange-brown, broadly oval in dorsal view, pars cephalica strongly elevated in lateral view, surface of elevated portion of pars cephalica granulate, sides granulate, with radiating rows of pits; lateral margin without denticles. Clypeus straight in front view, high, ALE separated from edge of carapace by their radius or more, around ALE diameter. Eyes relatively large, all subequal; ALE separated by more than their diameter, PLE-PME separated by less than PME radius. Sternum orange-brown, fused to carapace, furrow with rows of small pits, surface finely punctuate, microsculpture medially and in furrows, anterior margin with semicircular depression in the middle half, posterior margin not extending posteriorly of coxae IV; setae originating from surface. Mouthparts: Chelicerae, endites and labium orange-brown. Chelicerae anterior face unmodified, two pairs of medial strong setae; retromargin without teeth; fangs without tooth-like projections; paturon inner margin with pairs of enlarged setae, laminate groove absent. Labium anterior margin indented at middle. Endites anteromedian part forming a groove between serrula and the apical portion. ABDOMEN: pointed posteriorly; dorsum soft portions pale white. Book lung covers slightly elevated. Pedicel scuto-pedicel region with W-shaped scutal ridge, plumose setae dorsal and lateral of pedicel, scutal ridge medium sized with two strong downwards projected denticles. Dorsal scutum orange-brown, middle surface granulate, sides granulate. Epigastric scutum anterior spiracles secondary openings in an indent forming independent apertures. Postepigastric scutum orange-brown, almost semicircular, fused to epigastric scutum, without posteriorly directed lateral apodemes. Spinneret scutum large. LEGS: orange-brown. GENITALIA: Epigastric region with sperm pore circular, situated in front of anterior spiracles, rebordered. Palp proximal segments orange-brown. Conductor gently narrowing towards end with subdistal constriction, distal tip long, narrow and sharp pointed; embolar tip with short translucent embolar distal process; embolar prolateral process with narrow sclerotized area and broad translucent dorsal area ( Figs 49H , 50 A-D).

Female ( PBI _OON 31123). As in male except as noted. Total length: 1.64. GENITALIA: internal genitalia general shape subquadrate, with very short lateral processes, receptaculum rounded ( Fig. 51F ).

Other material examined. BRAZIL: Mato Grosso: Corumbá , -19.00960°, -57.65474°, May 05-08, 1964, B. Malkin, 1♂ ( AMNH PBI _ OON 01166) .

Distribution. This species is know from Costa Rica and Brazil.

Figures

FIGURES 31A–H. Cinetomorpha puberula Simon, male syntype (PBI_OON 05528). A, dorsal view. B, ventral view. C, lateral view. D, carapace, anterior view. E, sternum, ventral view. F, same, in clove oil immersion. G, abdomen, ventral view. H, same, in clove oil immersion. Abbreviations: Abbreviations: AS, anterior spiracle; G, gonopore; PS, posterior spiracle; SO anterior spiracle secondary opening.

FIGURES 32A–H. Cinetomorpha puberula Simon, male, syntype, SEM images. A-F, (PBI_OON 05528). G-H. (PBI_OON 26056).A, abdomen, lateral view. B, same, anterior view. C, G, palp, prolateral view. D, same, retrolateral view. E, H, same, anterior view. F, dorsal view.

FIGURES 33A–H. Cinetomorpha puberula Simon, male syntype. A-B, E-H, SEM images (PBI_OON 26056). C, D, transmitted light images (PBI_OON 05528). A, embolus, prolateral view. B, same, retrolateral view. C, same, dorsal view. D, epigastric area, ventral view. E, carapace, oblique lateral view. F, sternum, ventro-lateral view. G, endites and chelicerae, lateral view. H, chelicerae, antero-lateral view. Abbreviations: AS, anterior spiracle; C, conductor; E, embolus; EDP, embolar distal process; EPP, embolar prolateral process; ET, embolar tip; G, gonopore; PS, posterior spiracle; S, spermatic duct tip; SO anterior spiracle secondary opening.

FIGURES 42A–H. Cinetomorpha baja sp. nov., female. A-G, (PBI_OON 01182). H, (PBI_OON 02984). A, dorsal view. B, ventral view. C, carapace, dorsal view. D, lateral view. E, anterior view. F, sternum, ventral view. G, abdomen, anterior view. H, internal genitalia, ventral view, transmitted light image, clove oil immersion.

FIGURES 49A–H. Cinetomorpha longisetosa sp. nov., male (PBI_OON 31121). A, dorsal view. B, ventral view. C, carapace, dorsal view. D, lateral view. E, anterior view. F, sternum, ventral view. G, abdomen, anterior view. H, palp, dorso-prolateral view, transmitted light image, clove oil immersion.

FIGURES 50A–D. Cinetomorpha longisetosa sp. nov., male palp (PBI_OON 31121). A, prolateral view. B, retrolateral view. C, dorsal view. D, anterior view.

FIGURES 51A–F. Cinetomorpha longisetosa sp. nov., female (PBI_OON 31123).A, dorsal view. B, ventral view. C, carapace, anterior view. D, lateral view. E, abdomen, anterior view. F, internal genitalia, ventral view, transmitted light image, clove oil immersion.

FIGURES 52A–H. Cinetomorpha loreto sp. nov., male (PBI_OON 38819). A, dorsal view. B, ventral view. C, carapace, dorsal view. D, lateral view. E, anterior view. F, sternum, ventral view. G, abdomen, anterior view. H, palp, dorso-prolateral view, transmitted light image, clove oil immersion.

FIGURES 54A–H. Cinetomorpha loreto sp. nov., female (PBI_OON 38821). A, dorsal view. B, ventral view. C, carapace, dorsal view. D, lateral view. E, anterior view. F, sternum, ventral view. G, abdomen, anterior view. H, internal genitalia, ventral view, transmitted light image, clove oil immersion.

FIGURES 60A–H. Cinetomorpha zero sp. nov., male. A-G, (PBI_OON 38820). H, (PBI_OON 21098). A, dorsal view. B, ventral view. C, carapace, dorsal view. D, lateral view. E, anterior view. F, sternum, ventral view. G, abdomen, anterior view. H, bulb, dorso-prolateral view, transmitted light image, clove oil immersion.

FIGURES 63A–I. Cinetomorpha zero sp. nov., female A-H, (PBI_OON 21098). I, (PBI_OON 00018). A, dorsal view. B, ventral view. C, anterior view. D, carapace, lateral view. E, abdomen, lateral view. F, carapace dorsal view. G, sternum, ventral view. H, abdomen, anterior view. I, internal genitalia, ventral view, transmitted light image, clove oil immersion.