Pterostichus hoanglien Fedorenko, 2019

Fedorenko, D. N., 2019, New species of Pterostichus subgenus Circinatus (Coleoptera: Carabidae: Pterostichini) from Vietnam, Russian Entomological Journal 28 (4), pp. 358-369 : 368

5. Pterostichus hoanglien Fedorenko , sp.n.

Figs 5 , 10–11 , 19 , 28–30 , 39–40 .

MATERIAL. Holotype ♂ ( ZMMU) and paratypes ( SIEE), 8♂♂, 5♀♀, labelled: ‘N[orthern]- Vietnam, 40 km WNW of Lao Cai, env. Y Ty, Bat Xat Natn. Park, h= 1600–1900 m, 22°37´–37.6´N/ 103°37.3´–38.3´E, 16–21.X.2018, leg. D. Fedorenko’; 2♀♀, same data, except for 22°36´31´´N / 103°37´23´´E, h= 2000–2100 m, 4– 14.VI.2019; 2♀♀, same data except 22°37´36´´N / 103°37´32´´E, h= 1850 m.

DIAGNOSIS. As for the previous species, except that the body is slightly more robust, elytra much more rounded at humeri than behind, with apices truncate combined (i.e., angulate at sutural angles), elytral microsculpture consisting of transverse and very narrow meshes, mesepisterna invisible in dorsal view, pronotum with sublateral line indistinct or hardly traceable, and aedeagus distinctive. For other differences see the description.

DESCRIPTION. As for P. phangxipang sp.n. except as follows: Body ( Fig. 5 ), BL 10.4–12 mm. Head: Frontal sulci mostly entire and moderately deep.

Pronotum barely shorter and wider relative to head, PW/ PL 1.12–1.16 (mean 1.14, n=5), PW/HW 1.48–1.52 (1.50), with sides subconvex to indistinctly sinuate in basal third, and base barely wider, PB/PA 0.82–0.88 (0.85). Apical angles more produced due to apex truncate in between. Disc with fine yet distinct line running inside and parallel to lateral groove; this line being deeper in apical three fifths, very shallow to nearly indistinct behind. Basal sulci slightly short- er, running in basal third. Lateral bead fine and basally extended inside basal sulci.

Elytra barely shorter in general, EW/EL 1.54–1.58 (1.56), EW/PW 1.16–1.22 (1.18), with a nearly indistinct humeral tooth. Parascutellar striole missing. Discal seta d2 1/2–3/5 distant from base. USS: 15–16.

Prosternal process less blunt in lateral view; declivity flat in at least dorsal half.

Aedeagus ( Figs 10–11 , 19 , 28–30 ): median lobe sharply carinate in basal half of ventral side, with left margin cut in a straight line up to apex; apex rounded at right margin, without preapical tooth or dorsal groove. Right paramere triangular, slightly convex on right side, with thin yet not membranous integument on the reverse. Everted and inflated internal sac ( Figs 39–40 ) dorsal and spiral.

NAME. Toponymic noun, refers to Hoang Lien Mountain Ridge.

DISTRIBUTION. Known from the type locality only, the Bat Xat National Park , northern Vietnam .

HABITATS AND HABITS. The adults were taken under various rotten tree trunks and branches on the ground in cloudy broad-leaved or mixed forests, sometimes under bark of large standing dead trees, at height of more than 2 m above ground. The fact that the adults of both sexes were frequent in autumn (14 specimens in total), with only four females taken early in summer, may suggest the reproduction period in autumn or in late summer.

Figures

Figs 4–7. Dorsal habitus: 4 — Pterostichus phangxipang sp.n., female paratype; 5 — P. hoanglien sp.n., female paratype; 6 — Pterostichus batxatensis sp.n., holotype; 7 — P. laocaiensis sp.n., male paratype. Рис. 4–7. Габитус дорЗально: 4 — Pterostichus phangxipang sp.n., паратип самка; 5 — P. hoanglien sp.n., паратип самка; 6 — Pterostichus batxatensis sp.n., голотип; 7 — P. laocaiensis sp.n., паратип самец.

Figs 8–15. Median lobe of aedeagus: 8–9 — Pterostichus phangxipang sp.n.; 10–11 — P. hoanglien sp.n.; 12–13 — P. batxatensis sp.n.; 14–15 — P. laocaiensis sp.n.; 8, 10, 12, 14 — left view; 9, 11, 13, 15 — right view. Scale bars: 1 mm. Рис. 8–15. СреднЯЯ долЯ Эдеагуса: 8–9 — Pterostichus phangxipang sp.n.; 10–11 — P. hoanglien sp.n.; 12–13 — P. batxatensis sp.n.; 14–15 — P. laocaiensis sp.n.; 8, 10, 12, 14 — слева; 9, 11, 13, 15 — справа. Масштаб: 1 мм.

Figs 16–20. Median lobe of aedeagus: 16 — Pterostichus phangxipang sp.n.; 17–18 — P. batxatensis sp.n.; 19 — P. hoanglien sp.n.; 20 — P. laocaiensis sp.n.; 16–17, 20 — dorsal view; 18–19 — ventral view. Scale bar: 1 mm. Рис. 45–58. СреднЯЯ долЯ Эдеагуса: 16 — Pterostichus phangxipang sp.n.; 17–18 — P. batxatensis sp.n.; 19 — P. hoanglien sp.n.; 20 — P. laocaiensis sp.n.; 16–17, 20 — дорЗально; 18–19 — вентрально. Масштаб: 1 мм.

Figs 21–30. Right paramere: 21–22 — Pterostichus phangxipang sp.n.; 23–24 — P. batxatensis sp.n.; 25 — P. phiaoc sp.n.; 26–27 — P. laocaiensis sp.n.; 28–30 — P. hoanglien sp.n.; 21, 23, 25, 26, 28–29 — left view; 22, 24, 27, 30 — ventral view. Scale bar: 1 mm. Рис. 21–30. ПраваЯ парамера: 21–22 — Pterostichus phangxipang sp.n.; 23–24 — P. batxatensis sp.n.; 25 — P. phiaoc sp.n.; 26– 27 — P. laocaiensis sp.n.; 28–30 — P. hoanglien sp.n.; 21, 23, 25, 26, 28–29 — слева; 22, 24, 27, 30 — вентрально. Масштаб: 1 мм.

Figs 37–42. Everted and inflated internal sac of aedeagus: 37–38 — Pterostichus phangxipang sp.n.; 39–40 — P. hoanglien sp.n.; 41– 42 — P. laocaiensis sp.n.; 37, 41 — left view; 38 — dorso-apical view; 39 — dorsal view; 40 — left dorso-apical view; 42 — left ventrolateral view. Scale bar: 1 mm. Рис. 37–42. Вывернутый и надутый внутренний мешок Эдеагуса: 37–38 — Pterostichus phangxipang sp.n.; 39–40 — P. hoanglien sp.n.; 41–42 — P. laocaiensis sp.n.; 37, 41 — слева; 38 — дорЗоапикально; 39 — дорЗально; 40 — дорЗоапикально слева; 42 — вентрально слева. Масштаб: 1 мм.