Phylloclusia Hendel

Sasakawa, Mitsuhiro, 2011, Oriental species of clusiid flies (Diptera: Clusiidae: Clusiinae), Zootaxa 3038, pp. 1-28 : 2

Genus Phylloclusia Hendel

Phylloclusia Hendel, 1913: 78 . Type species, Phylloclusia steleocera Hendel, 1913 , fixed by monotypy.

Diagnosis. This genus is characterized by a large, triangular first antennal flagellomere with a bluntly angled dorsal apex (rarely small and rounded), the arista flattened in the form of a willow-leaf (rarely slender), the presence of prs and prsc, and the absence of if, presutural dc, and dp. Other diagnostic characters are as follows: occiput with shallow notch just behind ocellar triangle; distinct microtrichiation on wing membrane; fore femur with several fb; S6 setose posteriorly ( Lonsdale and Marshall 2007b); male cerci ( Figs 1, 2 , C) united at their bases and connected with posteroventral corners of epandrium, bearing many long setae posteriorly; surstylus ( Figs 1, 2 , S) varying in shape among species, setulose; proctiger ( Fig. 2 , P) Y-shaped; epiphallus distinct, mesophallus with sclerites separated or fused at their bases, distiphallus tubular, paraphallus short and pointed apically.

Remarks. Five Oriental species are known ( Hendel 1913; Sasakawa 1974; Lonsdale and Marshall 2007b), and two new species are described below.

Figures

FIGURES 1 – 5. Male genitalia of Phylloclusia bivittata sp. nov. (1 – 3, paratype) and P. sternopleuralis sp. nov. (4, 5, paratype). 1, 2, 4, 5, epandrium (E), surstylus (S), cercus (C), and proctiger (P), lateral view (1,4) and anterior view (2,5); 3, 4, hypandrium (H), phallapodeme (PA), and phallus, lateral view (parts of phallus: BP, basiphallus; DP, distiphallus; EA, ejaculatory apodeme; EP, epiphallus; MP, mesophallus; PP, paraphallus). Scale = 0.1 mm.