Neobarombiella senegalensis ( Laboissière, 1923 ) Bolz & Wagner, 2012

Bolz, Helmut & Wagner, Thomas, 2012, 3463, Zootaxa 3463, pp. 1-112 : 58-64

Neobarombiella senegalensis ( Laboissière, 1923) comb. n.

( Figs 92–105)

Barombia senegalensis Laboissière, 1923: 188 .

Barombiella leopoldi Laboissière, 1929: 153 syn. n.

Barombiella clathrata Laboissiere, 1939: 158 syn. n.

Barombiella costai Laboissière, 1939: 159 syn. n.

Barombia bicincta Laboissière, 1940: 81 syn. n.

Barombiella mutabilis Laboissière, 1940: 80 syn. n.

Barombiella pallida Laboissière, 1940: 80 syn. n.

Barombiella variabilis Laboissière, 1940: 87 syn. n.

Barombia basalis Laboissière , in litteris

Barombia crassicornis Laboissière , in litteris

Total length. 4.35–5.80 mm (mean: 5.01 mm) (n=42).

Head. Frons, vertex, labrum, labial and maxillary palpi yellow to yellowish-brown in about 75% of specimens studied, rest brown or black. Antenna pale yellow to yellow, apical antennomere dark brown or black ( Fig. 92); length of second to third antennomere 0.71–0.86 (mean: 0.77), and length of third to fourth antennomere 0.47–0.54 (mean: 0.50; Fig. 94). Eyes small, width of eye to interocular distance 0.42–0.48 (mean: 0.46) ( Fig. 92).

Thorax. Pronotum usually yellow or yellowish-brown, reddish-brown in 10% of specimens studied or very rarely black, as in 2% of specimens studied; very broad and strongly trapezoidal; pronotal width 1.68–2.06 mm (mean: 1.87 mm), pronotal length 0.80–0.94 mm (mean: 0.85 mm), and pronotal length to width 0.41–0.48 (mean: 0.45). Elytron entirely yellow in 10% of specimens studied) ( Fig. 92a) or yellow with shadow-like yellowish-red colouring on the elytral disc, as in lectotype of Barombiella pallida ( Fig. 102). Elytron usually pale yellow to yellow and black basally, which may be reduced to spot on either humerus, and transverse black band in apical third ( Fig. 92c); latter can be also reduced to transverse spots ( Fig. 99b); about 15% of specimens studied have red to brownish-red elytron with broad black base and broad sub-apical transverse black band, interrupted at suture, as in holotype of B. variabilis ( Fig. 103); very rarely with predominantly black elytron and yellow spots, as in holotype of B. leopoldi ( Figs 92g, 97); elytron broad, elytral length 3.60–4.40 mm (mean: 3.93 mm), elytral width 2.40–3.10 mm (mean: 2.69 mm), and maximal width of both elytra to length of elytron 0.65–0.74 (mean: 0.69). Meso- and metathorax yellow or yellowish-brown, legs yellow or entirely yellowish-brown, and length of basimetatarsus to metatibia 0.49–0.54 (mean: 0.52).

Abdomen. Yellow to yellowish-brown.

Male genitalia. Median lobe short and broad ( Fig. 93); sclerotized apex narrowing distinctly in lateral view with single pair of sclerotized ridges in apical quarter; deeply incised apically; endophallic brush with two slender spiculae is covered by tectum and median lobe.

Diagnosis. Neobarombiella flavilabris occurs sympatrically over the entire range of N. senegalensis and is the most similar in appearance to the latter, it also displays a similar highly variable colour pattern. In both species the polychroism has caused many synonymies, and since different colour patterns even occur syntopically, we found three junior synonyms of N. senegalensis described from the same locality (Rutshuru, Kivu). Both species can be reliably distinguished by the basal antennomeres, which are shorter and broader in N. senegalensis , length of third to fourth antennomere 0.47–0.54 in N. senegalensis (0.62–0.73in N. flavilabris ) ( Figs 9, 94);, N. senegalensis has very fine irregular dorsal punctuation, while that of N. flavilabris is usually much coarser, particularly on the elytra. However, due to the extreme variability in external characters in these two species, several doubtful cases may remain if based on external characters alone, The dissection of the male genitalia will allow reliable differentiation for both species ( Figs 8 and 93, and see the diagnosis for N. flavilabris ). Neobarombiella senegalensis occurs mainly in the Guineo-Congolian vegetation zone, and appears to be associated with forests, whereas N. flavilabris is particularly abundant in savannah habitats ( Figs 11, 95). Neobarombiella spielbergi sp. n. is similar to N. senegalensis in appearance, but its median lobe is far more slender, more elongate and rounded apically ( Figs 93, 186); and its third antennomere is more than half as long as the fourth antennomere (third antennomere only half as long as the fourth in N. senegalensis ) ( Figs 94, 187). Neobarombiella frontalis sp. n. and N. nigrosuturalis sp. n. can be distinguished from N. senegalensis by their colouring. Both species have black sutural margins. Neobarombiella frontalis sp. n. also has a prominent black spot on its vertex, which N. senegalensis lacks (Figs. 92,153). Neobarombiella cornuta sp. n. can be recognized by its longer pronotum, pronotal length to width 0.49–0.53 (0.41–0.48 in N. senegalensis ) ( Figs 92, 137). Neobarombiella flavicollis comb. n., N. medvedevi sp. n. and N. naumanni sp. n. can be characterised by their short second antennomere, length of second to third antennomere 0.60–0.70 in N. flavicollis ; 0.62–0.70 in N. naumanni sp. n.; and 0.58–0.64 in N. medvedevi sp. n. (0.71–0.86 in N. senegalensis ) ( Figs 84, 94, 167, 171).

Distribution. Recorded mainly in the Guineo-Congolian forest region, but occurs as far west as the Ivory Coast to Ethiopia and Kenya in the East and as far south as KwaZulu-Natal in South Africa, and is obviously quite abundant ( Fig. 95).

Type material examined. Barombia senegalensis . Lectotype, male: “Haut Sénégal, Khayes, Dr. Nodier., 6–8.1882 / Barombia , senegalensis, m., V. Laboissière – Dét. / Le Moult vend., via Reinbek, Eing Nr 1, 1957 / Type / AfriGa, specimen ID:, 1244, specimen data, documented, 26.IV.2006 / Neobarombiella , senegalensis (Lab. 1923) , Bolz & Wagner det. 2008 / Lectotypus, Th. Wagner desig. 2008, Barombia , senegalensis Lab. “ ( ZMUH; Fig. 96). All five syntypes mentioned by Laboissière in his original description are available. A lectotype is here designated. Type locality: Senegal, Khayes, 14°00'/ 14°00'W .— Paralectotypes: the four remaining specimens with the same label data as the lectotype are paralectotypes (3 ex. MNHN, 1 ex. ZMUH) .

Barombiella leopoldi . Holotype, female: “ Type / Musée du Congo, Env. Lac. Leopold II, 11–24–VI–1925, S. A. R. Prince Léopold / Barombia, Leopoldi , m, V. Laboissière – Dét., 1927 / R. Dét., 1814, W / Type / Holotypus, Barombia , leopoldi Lab., 1929 / Neobarombiella , senegalensis ( Laboissière, 1923) , Bolz & Wagner det. 2008 / AfriGa, specimen ID:, 1222, specimen data, documented, 5.VII.2005 ” ( MRAC; Fig. 97) . Holotype by monotypy “une female”. Colouring type: Fig. 92c. Type locality: Democratic Republic of the Congo, Lac Leopold, 1°04'S / 17°09'E .

Barombiella clathrata . Lectotype, female: “Elende, nov. / Angola, Miss. sc. suisse, 1932–1933 / Type / Le Moult vend., via Reinbek, Eing. Nr 1. 1957 / Barombiella , clathrata, m., V. Laboissière – Dét. / Lectotypus, Th. Wagner desig. 2008, Barombiella clathrata Lab. / Neobarombiella , senegalensis ( Laboissière, 1923) , Bolz & Wagner det. 04 / AfriGa, specimen ID:, 1208, specimen data, documented, 21.X.2005 ” (some cited labels not included in figure; ZMUH; Fig. 98). According to the original description five specimens were collected in Bimbi and Elende in Angola. Two of these specimens are available, and a lectotype is here designated. Colouring type Fig. 92g. Type locality: Angola, Elende, 12°44'S / 15°9'E .— Paralectotypes: the second specimen with the same label data as the lectotype is a paralectotype ( ZMUH) .

Barombiella costai . Lectotype, female: “Elende, nov. / Angola, Miss. sc. suisse, 1932–1933 / Cotype / Barombiella , costai m., V. Laboissière – Dét. / Lectotypus Th. Wagner desig. 2008, Barombiella , costai Lab. 1939 / Neobarombiella , senegalensis (Lab. 1923) , Bolz & Wagner det. 2008 / AfriGa, specimen ID:, 1211, specimen data, documented, 21.X.2005 ” ( ZMUH; Fig. 99). Of the seven specimens collected in November in Elende , which are listed in the original description, two type specimens are available and a lectotype is here designated. Colouring type Fig. 92d, but with reddish pronotum. Type locality: Angola, Elende, 12°44'S / 15°9'E .— Paralectotypes: the second specimen with the same label data as the lectotype is a paralectotype ( ZMUH) .

Barombia bicincta . Holotype, male: “ Type / Musée du Congo, Ruhengeri (source), 6–II–1935, G. F. de Witte, Parc Nat. Albert, 1069 / Barombia , bicincta, m., V. Laboissière – Det. / R. Dét., AA, 4572 / Type, B. bicincta / Holotypus, Barombia , bicincta Lab., 1940 / Neobarombiella , senegalensis (Lab., 1923) , Bolz & Wagner det. 2008 / AfriGa, specimen ID:, 1206, specimen data, documented, 5.VII.2005 ” ( MRAC; Fig. 100) . Holotype by monotypy “ 1 spécimen ”. Colouring type Fig. 92g. Type locality: Rwanda, Ruhengeri, 1°30’S / 29°38’E .

Barombiella mutabilis . Lectotype, male: “Type / Musée du Congo, Rutshuru, – V.1937, J. Ghesquière, 4521 / Barombiella , mutabilis m., V. Laboissière—Dét. / Type, mutabilis / R. Dét., 3922 / Lectotypus, Th. Wagner desig. 2005, Barombiella , mutabilis Lab. / Neobarombiella , senegalensis (Lab. 1923) , Bolz & Wagner det. 08 / AfriGa, specimen ID:, 1228, specimen data, documented, 5.VII.2005 ” ( MRAC; Fig. 101). Laboissière mentioned seven specimens in his original description and an undisclosed number of “variétés” from several locations which are regarded as syntypes. A lectotype is here designated. Colouring type Fig. 92c, but with reddish pronotum. Type locality: Democratic Republic of the Congo, Rutshuru, 1°11’S / 29°27’E .— Paralectotypes: 16 specimens with the same label data as the lectotype are paralectotypes (1 ex. BMNH, 2 ex. IRSN, 13 ex. MRAC) .

Barombiella pallida . Lectotype, male: “ Holotypus / Musée du Congo, Rutshuru, V–1937, J. Ghesquière, cafúin / Barombiella , pallida m, V. Laboissière. dét. 1939, Type. / R. Dét., A, 3679 / Lectotypus Barombiella pallida Lab., 1940 / Neobarombiella , senegalensis ( Laboissière, 1923) , Bolz & Wagner det. 2008 / AfriGa, specimen ID:, 1234, specimen data, documented, 5.VII.2005 ” ( MRAC; Fig. 102). Laboissière listed a total of 15 specimens from three different localities in Kivu in his original description. A lectotype is here designated. Colouring type Fig. 92a. Type locality: Democratic Republic of the Congo, Rutshuru, 1°11’S / 29°27’E.

Paralectotypes: nine specimens with the same label data as the lectotype (1 ex. BMNH, 8 ex. MRAC) ; 2 ex., Kivu Sake , V.1937 , Ghesquiere ( MRAC) ; 1 ex., Bombona , VII.1935 , A. Bai ( MRAC); two paralectotypes belong to N. flavilabris .

Barombiella variabilis . Holotype, male: “Type, variabilis / Musée du Congo, Rutshuru, V–1937, J. Ghesquière, 4521 / Type / R. Dét., W, 3921 / Barombiella , variabilis m., V. Laboissière—Dét. / Holotypus, Barombiella , variabilis Lab., 1940 / Neobarombiella , senegalensis (Lab. 1923) , Bolz & Wagner det. 2008 / AfriGa, specimen ID:, 1250, specimen data, documented, 5.VII.2005 “ ( MRAC; Fig. 103). The specimen from Rutshuru is designated as “type” in the original description. Colouring type Fig. 92f but with reddish elytra. Type locality: Democratic Republic of the Congo, Rutshuru, 1°11’S / 29°27’E .— Paratype: a single specimen with the same label data as the holotype is a paratype ( MRAC) .

Further material with type labels, but unpublished. Barombia basalis . 2 females: “Museum Paris, Ouganda, Onyoro Méridional, Ch. Alluaud 1909 / Janvier / Type / Barombia , basalis Labois, V. Laboissière – Dét. / Neobarombiella , senegalensis (Lab., 1923) , Bolz & Wagner 2008)” ( MNHN; Fig. 104). To our knowledge, a description of this species name has not been published, and it is hence unavailable within the meaning of the Code ( ZMUH) .

Barombia crassicornis . male: “Museum Paris, Afrique Orient. Angl., Boura, (Wa-Taita), Ch. Alluaud 1904 / Mars / Type / Barombia , crassicornis Labois, V. Laboissière —Dét. / Neobarombiella , senegalensis (Lab. 1923) , Bolz & Wagner det. 08” ( MNHN; Fig. 105). To our knowledge, a description of this species name has not been published, and it is hence unavailable within the meaning of the Code.

Other material examined. Burundi. 2 ex., Bururi, 3°58'S / 29°35'E, R . P. Giraudin ( MRAC); 2 ex., Bururi, 3°58'S / 29°35'E, III .1953, P. Basilewsky ( MRAC); 7 ex., Bururi, 3°58'S / 29°35'E, R . P. Giraudin ( MRAC) .— Cameroon. 1 ex., Jaunde Stat., 9°09'N / 14°17'E, Zenker S. ( MNHU); 1 ex., Joko , 5°29'N / 12°19'E, Staudinger ( ZMUH) .— Chad. 1 ex., Fort Sibut ( ZMUH) .— Democratic Republic of the Congo. 1 ex., Haut-Uele , Abimva, VIII .1925, L. Burgeon ( MRAC); 1 ex., Albertville , 5°56'S / 29°12'E, XII .1918, R. Mayne ( MRAC); 1 ex., Uele, Bambesa, 3°28'N / 25°43'E, IX.1933, Lefèvre ( MRAC); 2 ex., Uele , Bambesa , 3°28'N / 25°43'E, IX.1933, J. Leroy ( MRAC); 5 ex., Uelé , Bambesa , 3°28'N / 25°43'E, IX.1933 - X.1935, J. V . Leroy ( MRAC); 11 ex., Bambesa, 3°28'N / 25°43'E, IX.1932 – X.1938, J. Vrydagh ( MRAC); 24 ex., Bambesa, 3°28'N / 25°43'E, IX.1937 – I.1940, J. Vrydagh ( IRSN); 1 ex., Baraka , 4°20'N / 18°36'E, VIII .1918, R. Mayne ( MRAC); 1 ex., Beni , 0°30'S / 29°28'E, VIII .1932, L. Burgeon ( MRAC); 1 ex., Blukwa, 1°45'N / 30°36'E, XI.1928, A. Collart ( IRSN); 1 ex., Bomboma , 1°14'N / 23°58'E, VII .1935, A. Bai ( MRAC); 2 ex., Ituri, Bunia, 1°34'N / 30°15'E, 1938, P. Lefèvre ( MRAC); 5 ex., Congo da Lemba , 5°42'S / 13°42'E, I.–II.1913, R . Mayne ( MRAC); 1 ex., Dele, 1°32'E / 30°16'E, IV.1935, H. J. Bredo ( MRAC); 4 ex., Uele, Dingila , 3°39'N / 26°04'E, VIII .1933, H. J. Bredo ( MRAC); 7 ex., Uele, Dingila , 3°39'N / 26°04'E, V .– VII.1933, J. V. Leroy ( MRAC); 1 ex., Eala, 0°04'N / 18°17'E, I.1936, J. Ghesquière ( MRAC); 1 ex., Elisabethville, 11°40'S / 27°28'E, 1939, H. J. Brédo ( IRSN); 2 ex., Elisabethville, 11°40'S / 27°28'E, II.1952, Ch. Seydel ( MRAC); 1 ex., Fort Crampel, I.1957 ( ZMUH); 1 ex., Gandajika , 6°45'S / 23°56'E, XII .1950, P. de Francquen ( MRAC); 31 ex., Kivu, Ibanda, 2°13'S / 17°01'E, 1952, M. Vandelannoite ( MRAC); 1 ex., P. N. Albert, Kabasha , 0°45'S / 29°13'E, VII .1934, G. F. de Witte ( MRAC); 1 ex., Lualaba, Kabongo, 4°07'S / 15°31'E, XI.1953, Ch. Seydel ( MRAC); 1 ex., Kamponde, 6°42'S / 22°56'E, 1945, Rèv. Fr. Allaer ( MRAC); 1 ex., P. N. Upemba, Kanonga, 9°20'S / 27°12'E, 1949, Miss. G. F. de Witte ( IRSN); 4 ex., Lulua, Kapanga, 8°21'S / 22°35'E, X.1932 – II.1933, F. G. Overlaet ( MRAC); 1 ex., Kasongo , 6°11'S / 27°34'E, III .1960, P. L. G. Benoit ( MRAC); 2 ex., Kaziba, 7°09'S / 27°01'E, II.1948, G. F. de Witte ( MRAC); 1 ex., Kikwit, 5°03'S / 18°52'E, 1920, P. Vanderijst ( MRAC); 1 ex., Kindu, 2°57'S / 25°57'E, 1917, L. Burgeon ( MNHN); 1 ex., Lomami, Kisamba, 9°07'S / 26°17'E, I.1931, P. Quarré ( MRAC); 1 ex., Kisantu, 5°07'S / 15°05'E, P. Goossens ( MRAC); 1 ex., Kolwezi , 10°43'S / 25°28'E, VIII .1961, Dr. V . Allard ( MRAC); 1 ex., Bangala, Kutu , 2°31'N / 19°25'E, VI .1935, G. Settembrino ( IRSN); 1 ex., Kwangila , 2°30'N / 27°37'E, XII .1923, A. Collart ( MRAC); 2 ex., Libenge , 3°39'N / 18°38'E, 1947, R . Cremers & M. Neuman ( IRSN); 1 ex., Kivu, Lulenga, 0°37'S / 18°23'E, IX.1932, L. Burgeon ( MRAC); 1 ex., M'Paka , 4°05'N / 19°13'E, XII .1959, M. Pecheur ( MRAC); 2 ex., Mahagi-Niarembe, 2°15'N / 31°07'E, 1935, Ch. Scops ( MRAC); 1 ex., Mai-Ndombe, 1°04'S / 17°09'E ( MRAC); 1 ex., Kivu , Malungu , 2°52'S / 27°07'E, IV.1935, J. V . Leroy ( MRAC); 4 ex., Kivu, Malungu, 2°52'S / 27°07'E, 1939, Hendrickx ( MRAC); 1 ex., Mandungu , 2°25'N / 23°13'E, XI.1912, R . Mayne ( MRAC); 3 ex., Mayidi, 5°11'S / 15°09'E, 1942, Rév. P. Van Eyen ( MRAC); 10 ex., P. N. Garamba, Miss. H. De Saeger , 3°40'N / 29°00'E, VII .1950, G. Demoulin ( IRSN); 2 ex., Modu , VI .1931, H. J. Bredo ( MRAC); 2 ex., Haut-Uele, Moto, 2°27'N / 26°25'E, IV.1925, L. Burgeon ( MRAC); 2 ex., Moyen , 1919, Pitard ( MNHN); 1 ex., Kivu , Mulungu , 2°20'S / 26°04'E, V .1954, J. Decelle ( MRAC); 3 ex., Kivu, Mulungu , 2°20'S / 26°04'E, VIII .1952, R. Mayne ( MRAC); 8 ex., Kivu, Mulungu, 2°20'S / 26°04'E, 1938-1949, Hendrickx ( MRAC); 2 ex., P. N. Albert, Mutsora , 2°10'N / 26°00'E, 1939, Hackars ( IRSN); 1 ex., Ngowa , 3°42'S / 16°35'E, IV.1939, J. Mertens ( IRSN); 1 ex., Niembo , 7°22'S / 25°52'E, VII .1918, R. Mayne ( MRAC); 3 ex., Kibali-Ituri, Nioka , 2°57'S / 26°10'E, 1933– VIII .1952, J. Hecq, Lefèvre, H. J. Bredo ( MRAC); 2 ex., P. N. Garamba, Pidigala, 4°35'N / 29°32'E, IV.1952, H. De Saeger ( MRAC); 1 ex., P. N. Albert, Rumaegabo, 2°10'N / 26°00'E, IV.1945, G. F. de Witte ( MRAC); 1 ex., pont Kako, Rutshuru, 1°11'S / 29°27'E, IX.1932, L. Burgeon ( MRAC); 59 ex., Rutshuru , 1°11'S / 29°27'E, V .1937, J. Ghesquière (1 ex. BMNH / 58 ex. MRAC); 2 ex., Kivu, Rwankwi , IV.1946 – VII .1951, J. V. Leroy ( MRAC); 4 ex., Kivu, Sake , 0°30'S / 28°41'E, V .1937, J. Ghesquière ( MRAC); 1 ex., Stanleyville, X.1925, J. Ghesquière ( MRAC); 1 ex., Stanleyville , VII .1928, A. Collart ( MRAC); 4 ex., Kivu, Tshamfu , 2°36'S / 28°50'E, VI .1938, L. Hendrickx ( MRAC); 2 ex., Tshuapa Bokungu, 0°41'S / 22°19'E, 1949, Dupuis ( MRAC); 16 ex., Uelé , X.1938, R . v. d. Veen ( RMNH); 2 ex., Uvira , 3°24'S / 29°08'E, XII .1932, L. Burgeon ( MRAC); 1 ex., Haut-Uele, Watsa, 0°33'S / 24°46'E, 1922, L. Burgeon ( MRAC); 1 ex., Kivu , IV.1966, M. J. Celis ( MRAC); 1 ex., P. N. Albert, 2°10'N / 26°00'E, I.1954, H. Synave ( IRSN); 1 ex., P. N. Upemba, 9°20'S / 27°12'E, II.1948, Mis. G. F. de Witte ( IRSN); 1 ex., Region des Lacs , Dr.Sagona ( MRAC); 1 ex., Tanganyika Terr., IV.1957, P. Basilewsky ( MRAC); 7 ex., Uele , 3°37'N / 28°33'E, X.1938, R . v. d. Veen ( RMNH) .— Ethiopia. 2 ex., Shou Provinz, Langano Lake , 7°35'N / 38°50'E, IX.1973, R . O. S. Clarke ( MRAC); 1 ex., Omo , 7°37'N / 37°15'E, V .1904, Besche & Hauasch ( ZISP) .— Equatorial Guinea. 2 ex., 02°00’N / 10°00’E, Dr. L. Báguena ( MNCN); 1 ex., Evinayong , 01°06’N / 10°17’E, Dr. L. Báguena ( MNCN) .— Ivory Coast. 1 ex., Eremankono , 5°33'N / 5°22'W, VIII .–IX.1962, J. Decelle ( MRAC); 1 ex., Kouihly, 8°00'N / 5°00'W, 1938, L. Chopard ( MNHN); 1 ex., Pays Dyola, Sassandra , 8°53'N / 7°52'W, 1910, A. Chevalier ( MNHN) .— Kenya. 1 ex., Mathews Range, Kichich , 1°12'N / 37°17'E, 1978, R . De Jong ( RMNH) .— Malawi. 1 ex., Fort Lister , XII .1983, Dept. Entomology ( SANC); 1 ex., Lilongwe, V .– VI .1910, S. A. Neave ( BMNH); 3 ex., Mlanje , 16°02'S / 35°30'E, I.– V .1913, S. A. Neave ( BMNH); 3 ex., Ruo Valley , 16°33'S / 35°09'E, XII .1913, S. A. Neave ( BMNH) .— Mozambique: 2 ex., Guengère , 19°04'S / 34°13'E, 1906, G. Vasse ( MNHN) .— Namibia. 1 ex., Kunene River, 17°24'S / 14°00'E, II.1975, Endrödy-Younga ( TMSA); 1 ex., Otjivarongo , 18°50'S / 14°22'E, XII .1949, G. Hobohm ( TMSA).— Nigeria. 1 ex., Lamu , 08°47’N / 12°28’E ( MNCN) .— Rwanda. 3 ex., Col. Kobayaza, 1°34'S / 29°32'E, XI.1951, A. E. Bertrand ( MRAC); 5 ex., Kagogo, I.1953, P. Basilewsky ( MRAC); 1 ex., Kayove, 1°53'S / 29°21'E, II.1953, P. Basilewsky ( MRAC); 1 ex., Kibungu , 2°06'S / 30°02'E, X.– XII .1937, R . Verhulst ( MRAC); 2 ex., Kinazi , 2°11'S / 29°55'E, I.1953, P. Basilewsky ( MRAC); 12 ex., Mahembe, 1°52'S / 29°31'E, I.1953, P. Basilewsky ( MRAC); 1 ex., Muhavura, I.1953, P. Basilewsky ( MRAC); 1 ex., Nyakabye, 2°34'S / 28°01'E, IV.1984, H. Mühle (Cki); 2 ex., Oyangugu, VI .1983, H. Mühle (Cki); 3 ex., Ruhengeri , 1°30'S / 29°38'E, VIII .–X.1934, G. F. de Witte ( MRAC); 2 ex., Ruhengeri , 1°30'S / 29°38'E, I.1953, P. Basilewsky ( MRAC) .— South Africa: South Africa: Mpumalanga. 2 ex., Pilgrims Rest , 24°54'S / 30°45'E, XII .1995, R. Stals ( SANC) .— Tanzania. 8 ex., Bugala, 6°17'S / 30°07'E, 1908, Dr. E. Bayon ( MCSG); 1 ex., Usambara, Nguelo , 8°40'S / 3°13'E ( MNHU); 1 ex., Tanganyika , 9°07'S / 35°38'E, XII .1935, Amani ( BMNH); 1 ex., Usambara , 4°45'S / 38°02'E, X.1931, S. Conradt ( MNHU) .— Togo. 1 ex., Bismarckburg, VI .1893, L. Conradt ( MNHU) .— Uganda. 1 ex., Mpanga Forest, 0°06'N / 32°00'E, XI.1911, S. A. Neave ( BMNH); 2 ex., Kibale Nat. Park, Fort Portal , 0°41'N / 30°15'E, VII .– VIII .1998, L. Schmidt ( ZFMK); 2 ex., Hoima, 1°26'N / 31°20'E, XII .1911, S. A. Neave ( BMNH); 2 ex., Kampala, 0°18'S / 32°33'E, XII .1932, A.F.J. Gedye ( BMNH); 2 ex., Kibale Forrest , 0°37'S / 30°16'E, II.1984 – III .1985, M. Nummelin ( MZHF); 4 ex., Mitanda , 0°36'S / 30°09'E, III .1932, H. Hargreaves ( BMNH) .— Zambia. 1 ex., Chirinba Forrest , 12°54'S / 32°6'E, V .1907, C. F. M. Swynnerton ( BMNH); 1 ex., Kashitu, 13°45'S / 28°40'E, XI.1914, H. C. Dollmann ( BMNH); 1 ex., Mumbwa , 14°59'S / 27°04'E, IX.1913, H. C. Dollmann ( BMNH) .— Zimbabwe. 1 ex., Mawhuradonha, XII .1998, J. Běcvář (CBez).