Neodima sichuanensis Schimmel & Platia, 1992

Kundrata, Robin, Sormova, Eliska, Qiu, Lu & Prosvirov, Alexander S., 2019, Revision of the flightless click-beetle genus Neodima Schimmel & Platia (Elateridae: Dimini) endemic to China, with comments on its systematic position, Zootaxa 4604 (1), pp. 42-58 : 42-58

publication ID	https://doi.org/ 10.11646/zootaxa.4604.1.2
publication LSID	lsid:zoobank.org:pub:42FB8004-F957-45E7-AA3C-58D47E1A8FDD
DOI	https://doi.org/10.5281/zenodo.5943252
persistent identifier	https://treatment.plazi.org/id/03A8878B-FF8E-FFF5-FF48-45168A8C8B76
treatment provided by	Plazi
scientific name	Neodima sichuanensis Schimmel & Platia, 1992
status

Treatment

( Figs 1 View FIGURE 1 A–C; 2; 3)

Neodima sichuanensis Schimmel & Platia, 1992: 300 ; Schimmel (1996: 154), Schimmel (1998: 223), Cate et al. (2007: 184), Kundrata et al. (2018b: 30).

Type material. Holotype, male ( PCCW); 7 paratypes: 1 male ( NMPC), 3 ex., sex undetermined ( PCCW), 1 female (coll. J. Mertlik, Czech Republic), 1 female (coll. V. Dušánek, Czech Republic), 1 ex., sex undetermined ( PCRS).

Material examined. Holotype, male, “ China, Sichuan, Mt. Emei , 600–1050 m, 5.– 19.5.1989, Lad. Bocák lgt. // Holotypus, Neodima n. gen. sichuanensis n. sp. det. Schimmel & Platia, 1990” ( PCCW) . Paratype, male, “ China, Sichuan, 103.20el, 29.30nw, Mt. Emei , 500–1200 m, 4.–19.V.1989, S. & J. Kolibáč leg.” ( NMPC) ; male, “ China, SW Sichuan, Mt. Emei, 6.6.[19]97, Krajcik M. lgt. // Czikia catei Schimmel & Platia, 1998, Vít Kubáň det., VI.1998 // Neodima sichuanensis Schi. , det. Schimmel “ ( NMPC) GoogleMaps ; 3 males, „P. R. China, Sichuan, Emei Shan, N29°34‘46‘‘, E103°22‘04‘‘, 1463 m, 27.V.2011, sift07, V. Grebennikov [lgt.]“ ( CNCI) GoogleMaps ; 1 female, “P. R. China, Sichuan, Emei Shan, N29°33’00’’, E103°21’38’’, 1639 m, 28.V.2011, sift08, V. Grebennikov [lgt.]“ ( CNCI) GoogleMaps ; 1 female, „[ China], Mt. Emei , Sichuan, VII. 2003 “ ( PCLQ) GoogleMaps .

Diagnosis. This species differs from its congeners by having the antennomere IV only 1.20–1.25 times longer than antennomere II (1.40–1.50 times in other species; Figs 2B View FIGURE 2 , 4B View FIGURE 4 , 6B View FIGURE 6 , 7A View FIGURE 7 ), concave frontal margin of mesanepisternum (straight in other species; Figs 2C View FIGURE 2 , 3D View FIGURE 3 , 4C View FIGURE 4 , 6C View FIGURE 6 , 7D View FIGURE 7 ), relatively short elytra (up to 1.75 times as long as wide; in other species 1.85–2.20 times; Figs 1A, D View FIGURE 1 ; 3A View FIGURE 3 ; 5A, D, G View FIGURE 5 ), and dark elytral stripes not reaching base of elytra. Except N. sichuanensis , male genitalia are known only for N. belousovi Prosvirov & Kundrata sp. nov. The latter species differs in having a median lobe distinctly surpassing parameres and having a large hook-like sclerite inside (only slightly surpassing parameres and without sclerites inside in N. sichuanensis ), and a simple paramere apex (with subapical hook in N. sichuanensis ) ( Figs 2 View FIGURE 2 I–K; 6L–N).

Redescription. Male. Body length 4.50–5.20 mm (holotype 4.75 mm); width 1.75–1.85 mm (holotype 1.75 mm). Body ( Fig. 1 View FIGURE 1 A–C) slightly elongate, light brown to reddish brown; antennomere apices, head, pronotal disc darker, usually dark brown, elytra yellowish to yellowish brown, each elytron with dark brown elongate stripe between stria III and stria VII running along almost entire elytral length, legs yellowish to light brown; pubescence yellowish to golden.

Head ( Fig. 2A, D View FIGURE 2 ) including eyes 0.60–0.65 times as wide as pronotum; supra-antennal carinae smooth, distinct, medially forming edge but not sharp frontal carina. Head surface sparsely and coarsely punctate; punctures large, suboval, intervals between punctures usually 1–2 times puncture diameter. Maxillary palpus with palpomere III slightly longer than wide. Antenna ( Figs 1 View FIGURE 1 A–C, 2B) slightly surpassing midlength of elytra; length ratio of antennomeres II–IV = 1.00: 1.15–1.20: 1.20–1.25; pedicel shortest, slightly more than twice as long as wide; antennomeres III–X elongate, about 2.5 times as long as wide; ultimate antennomere elongate, about 1.2 times as long as penultimate antennomere, about three times as long as wide, gradually widened subapically, obliquely narrowed apically.

Pronotum ( Fig. 2D View FIGURE 2 ) 1.15–1.25 as wide as long if measured along midline (length at midline 1.25–1.50 mm; width 1.45–1.75 mm), and 0.95–1.00 as wide as long if measured between anterior and posterior angles, strongly convex. Anterior angles slightly produced forward; lateral sides slightly rounded; posterior angles short, relatively sharp, very slightly divergent. Disc sparsely punctate; punctures large, moderately deep, mostly oval, separated usually 2–4 times of their diameter, near margins smaller; interstices almost smooth. Hypomeron moderately densely punctate, with punctures large, rounded to oval, distinct, separated usually 2–3 times of their diameter. Prosternum ( Figs 1B View FIGURE 1 , 2C View FIGURE 2 ) including prosternal process 1.6 times as long as wide, in front of coxal cavities 0.9 times as long as wide; with punctation similar to hypomeron or slightly denser; prosternal process about 0.7 times as long as prosternum in front of procoxal cavities. Scutellar shield ( Fig. 2E View FIGURE 2 ) suboval, about 1.3–1.4 times as wide as long; anterior margin slightly rounded, posterior margin rounded; surface covered with moderately dense small punctures. Mesoventrite ( Figs 1B View FIGURE 1 , 2C View FIGURE 2 ) transverse, medial prostrusion of frontal margin apically distinctly emarginate medially; procoxal rests indistinct. Mesanepisternum with frontal margin medially smoothly but distinctly emarginate, without apparent lateral extensions of procoxal rests. Elytra ( Fig. 1A, C View FIGURE 1 ) oval-elongate, together 0.60–0.65 times as long as body, 1.65–1.75 times as long as wide, widest at apical 2/3, sides slightly rounded; with striae formed by lines of window-like large punctures, smaller and more rounded near margins, larger and more rectangular medially; interstriae reduced to narrow smooth ridges; laterally gradually bent downwards after interstria VII at basal half.

Abdomen ( Fig. 1B View FIGURE 1 ) with ventrites moderately densely puncate, punctures large, oval; apical ventrite widely rounded apically. Sternite IX ( Fig. 2G View FIGURE 2 ) about twice as long as wide, with apex narrowly rounded, covered with short pubescence; tegite IX ( Fig. 2H View FIGURE 2 ) wider than long, deeply emarginate medially; tergite X ( Fig. 2H View FIGURE 2 ) small, elongate, narrowly rounded apically. Aedeagus ( Fig. 2 View FIGURE 2 I–K) about 2.8 times as long as wide. Median lobe elongate, slightly surpassing tip of parameres, from about half gradually narrowed towards apex, apically narrowly rounded, slightl curved from lateral view; with very short basal struts. Paramere elongate, relatively broad at basal half, then distinctly narrowed and before apex widened to form a distinct flattened subapical hook oriented laterally, apex subacute, area between apex and subapical hook straight. Phallobase relatively small, transverse, basally slightly emarginate medially.

Female ( Fig. 3 View FIGURE 3 ). Body length 5.30–5.75 mm; width 1.90–2.25 mm. Similar to male in almost all characters ( Fig. 3 View FIGURE 3 A–F); antenna slightly shorter, reaching about midlength of elytra, pronotum more convex, with posterior angles not so sharp and more divergent ( Fig. 3A, C View FIGURE 3 ). Tergite VIII ( Fig. 3G View FIGURE 3 ) broadly triangular, about 1.4 times as long as wide, apically gradually narrowed, finely punctate, covered with sparse pubescence. Sternite VIII ( Fig. 3G View FIGURE 3 ) triangular, about as long as wide, apically widely rounded, spiculum ventrale about five times sternite length. Ovipositor ( Fig. 3H View FIGURE 3 ) with paraprocts about five times as long as gonocoxites; gonocoxite apically narrowed; stylus elongate. Bursa copulatrix ( Fig. 3H View FIGURE 3 ) elongate, sac-like, without any sclerotized structures.