Tomosvaryella angulata Kehlmaier & Majnon-Jahromi, 2017

Motamedinia, Behnam, Skevington, Jeffrey H. & Kelso, Scott, 2021, Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species, Zootaxa 5002 (1), pp. 1-103 : 14-16

Tomosvaryella angulata Kehlmaier & Majnon-Jahromi, 2017

Figs 4A‒E , 50

Diagnosis: This species can be recognized by the shape of surstyli, both apically knobbed in dorsal view ( Fig. 4A ) and bent strictly towards the abdominal sternites in lateral view ( Fig. 4D‒E ); phallus trifid and unequal, one with tooth-like projections in basal half ( Fig. 4D‒E ); phallic guide short and broad ( Fig. 4D‒E ); gonopods slightly unequal in height, left is slightly higher than right ( Fig. 4B ). The female is described and illustrated in Majnon-Jahromi et al. (2017b).

Specimens examined: IRAN: Sistan & Baluchestan: Saravan , 27°25’N, 62°17’E, 12.v.2013, F. Hamzavi ,

Malaise trap, JSS51917 (1♂, CNC) ; Kermanshah : Sarpolezahab, 34°28’N, 45°49’E, 456 m a.s.l., 14.v.2016, M. Zardouei, Malaise trap, JSS52190 (1♂, CNC) ; Khuzestan : Shush, 32°06’N, 48°26’E, 68 m a.s.l., 11.iii–10.v.2015, E. Gilasian, Malaise trap, JSS52192 (1♀, CNC) ; UNITED ARAB EMIRATES: Barqa Al Suqoor Protected Area , 23°59’N, 52°40’E, 29.iii.2018, A. Saji & A. van Harten, Malaise trap, JSS52275 (1♂, CNC) .

Distribution: Iran, United Arab Emirates ( Skevington 2020) ( Fig. 50 ).

Note: Based on DNA barcoding, this species is most similar to T. pseudophanes ( Perkins, 1905) , differing by 8.3% (pairwise divergence). Intraspecific variation ranges from 0.1 to 3.2% (Supplementary Table 1).

Figures

FIGURE 4. Male genitalia of Tomosvaryella angulata (JSS52275, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view.

FIGURE 50. Distribution of Tomosvaryella angulata, T. argyrata, T. argyratoides, T. cyprusensis sp. nov. in the Middle East.