Copris remotus

Kohlmann, Bert & Delgado, Enio Cano And Leonardo, 2003, New species and records of Copris (Coleoptera: Scarabaeidae; Scarabaeinae) from Central America, Zootaxa 167, pp. 1-16 : 12

Key to the species of the remotus complex based on the major males

1. Head unarmed having only a gibbosity, clypeus tridentate; thorax unarmed. Costa Rica........................................................................................ tridentatus Solís & Kohlmann

­ Head armed with a horn, clypeus bidentate; thorax armed .......................................... 2

2. Pygidial margin incomplete, its inner border completely effaced ventrally. Chiapas, Guatemala, Costa Rica and Panamá.............................................. costaricencis Gahan

­ Pygidial margin complete, its inner border entirely engraved ...................................... 3

3. Eighth elytral stria incomplete; frontal pronotal declivity with a carina running through its middle ....................................................................................................... 4

­ Eighth elytral stria complete; frontal pronotal declivity without a carina running through its middle ....................................................................................................... 5

4. Antennae dark­brown; male anterolateral pronotal angles not forming median point, but presenting an excavation behind these angles ( Fig. 4 ), median pronotal prominences diverging ( Fig. 1 ). Guatemala. caliginosus Kohlmann, Cano & Delgado , n. sp.

­ Antennae reddish­brown; male anterolateral pronotal angles forming a small downwardly directed median point and separated from the angles by a more or less level area ( Fig. 4 ), median pronotal prominences strongly diverging ( Fig. 2 ). Guatemala and Honduras............................................. nubilosus Kohlmann, Cano and Delgado, n. sp.

5. Protibial apical spur abruptly bent inward very near apex. Oklahoma, Texas, Coahuila, Nuevo León and Tamaulipas .............................................................. remotus LeConte

­ Protibial apical spur curved or slightly curved inward ( Fig. 3) ................................... 6

6. Anterior pronotal margin with a minute, acute median tooth ( Fig. 3); elytral striae with large, less transverse punctures, distinctly crenating the margins of the strial channels; protibial apical spurs blunt ( Fig. 3). Veracruz to Chiapas ........................ sallei Harold

­ Anterior pronotal margin not forming any median point or angle; elytral striae with fine, very transverse punctures which at most only slightly crenate the edges of the channel; protibial apical spurs pointed. Michoacán .. mexicanus Matthews and Halffter

Figures

FIGURE. 1. Dorsal view of Copris caliginosus Kohlmann, Cano and Delgado, sp. nov. (holotype).

FIGURE. 2. Dorsal view of Copris nubilosus Kohlmann, Cano and Delgado, sp. nov. (holotype).

FIGURE. 4. Lateral views of head and pronotum of Copris caliginosus Kohlmann, Cano and Delgado, sp. nov. (holotype, top), Copris nubilosus Kohlmann, Cano and Delgado, sp. nov. (holotype, middle) and Copris sallei (bottom).