Raveniola fedotovi ( Charitonov, 1946 )

Zonstein, Sergei L., 2024, A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia, European Journal of Taxonomy 967, pp. 1-185 : 52-54

publication ID	https://doi.org/ 10.5852/ejt.2024.967.2699
publication LSID	lsid:zoobank.org:pub:C08B8027-50CC-417E-BCD4-5183B9FF6738
DOI	https://doi.org/10.5281/zenodo.14034706
persistent identifier	https://treatment.plazi.org/id/03A8B723-5519-FF8F-FDD9-E318FD84C9B2
treatment provided by	Plazi
scientific name	Raveniola fedotovi ( Charitonov, 1946 )
status

Treatment

Raveniola fedotovi ( Charitonov, 1946) View in CoL

Figs 16 View Figs 10–18 , 51 View Figs 45–53 , 98 View Figs 91–99 , 125 View Figs 118–126 , 153 View Figs 148–159 , 183 View Figs 172–183 , 199 View Figs 196–201 , 217 View Figs 211–219 , 244 View Figs 238–246 , 274–275 View Figs 274–281 , 366 View Figs 364–378 , 432–433 View Figs 429–438 , 528–529 View Figs 522–536 , 591–592 View Figs 584–592 , 707–710, 755

Brachythele fedotovi Charitonov, 1946: 19 , fig. 2 (♂).

Brachythele fedotovi – Charitonov 1969: 65. — Brignoli 1983: 123. — Zonstein 1985: 158.

Raveniola fedotovi View in CoL – Zonstein 1987: 1015. — Platnick 1989: 90. — Mikhailov 1996: 77; 1997: 20; 2013: 12. — Zonstein et al. 2018b: 66, fig. 100 (♂). — Zonstein & Esyunin 2023: 76, figs 1–14 (♂ ♀).

Diagnosis

Males of Raveniola fedotovi differ from other male congeners in having a characteristic triangular (very short, but high and acute) keel, demarcating proximal and distal sections of the embolus (vs either the presence of a lengthened keel in R. diluta sp. nov., or the absence of a raised keel in R. pallens sp. nov. and R. zyuzini sp. nov.; Figs 432–433 View Figs 429–438 , cf. Figs 428–431, 324–438 View Figs 418–428 View Figs 429–438 View Figs 319–333 View Figs 334–348 View Figs 349–363 View Figs 364–378 View Figs 379–388 View Figs 389–399 View Figs 400–408 View Figs 409–417 ). The sole adult female of R. fedotovi , known to date, is distinguishable owing to a specific structure of the spermathecal trunks, which are considerably longer, and relatively more closely spaced to each other than in females of other species representing the same group ( Figs 528–529 View Figs 522–536 , cf. Figs 526–527, 530–533 View Figs 522–536 ).

Material examined

Lectotype

UZBEKISTAN • ♂; Hissar Mts (northwestern slope) , surroundings of Ishkent Village; 38°49′ N, 66°58′ E; 1100–1300 m a.s.l.; 25–28 Mar. 1942; D.M. Fedotov leg.; ZMMU. GoogleMaps

Paralectotypes

UZBEKISTAN • 3 ♂♂; same collection data as for holotype; PSU, deposited as preparations GoogleMaps .

Additional material (1 ♂, 1 ♀, 2 juvs)

UZBEKISTAN • 1 juv.; Kugitang (Koitendagh) Mts , Baglydara Canyon , 10–11 km W of Hatak Village; 37°58′ N, 66°43′ E; 1300–1400 m a.s.l.; 8 Apr. 1989; S. Zonstein leg.; SMNH GoogleMaps • 1 juv.; Baisuntau Mts , Akrabat Pass ; 38°15′ N, 66°50′ E; 1500 m a.s.l.; 17 Apr. 1987; S. Zonstein leg.; SMNH GoogleMaps • 1 ♂; Zeravshan Mts , Jindy-Daria Canyon , Hojakurgan Gorge ; 39°11′ N, 67°17′ E; 1400–1600 m a.s.l.; 29 Apr. 1992; S. Zonstein leg.; SMNH GoogleMaps • 1 ♀; same collection data as for preceding; 4 May 2022; S. Zonstein leg.; SMNH GoogleMaps .

Description

Male (lectotype)

Figures 16 View Figs 10–18 , 153 View Figs 148–159 , 199 View Figs 196–201 , 217 View Figs 211–219 , 274–275 View Figs 274–281 , 366 View Figs 364–378 , 432–433 View Figs 429–438 , 591 View Figs 584–592 are based on the paralectotypes and conspecific material.

HABITUS. See Fig. 16. View Figs 10–18

MEASUREMENTS. TBL 6.90, CL 3.38, CW 2.85, LL 0.28, LW 0.60, SL 1.78, SW 1.51.

COLOUR. Carapace, palps and legs light brownish orange; eye tubercle blackish brown; chelicerae light reddish brown; sternum, labium and maxillae yellow; abdomen uniformly greyish white without clear dorsal pattern.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 98 View Figs 91–99 . Clypeus and eye group as in Fig. 153 View Figs 148–159 . Eye diameters and interdistances: AME 0.08(0.12), ALE 0.13, PLE 0.09, PME 0.06; AME–AME 0.08(0.05), ALE–AME 0.05(0.04), ALE–PLE 0.02, PLE–PME 0.02, PME–PME 0.16. Anterior cheliceral edge with unmodified setae; rastellum not developed. Each cheliceral furrow with 7 promarginal teeth and 4–5 mesobasal denticles. MIT indiscernible ( Fig. 199 View Figs 196–201 ). Sternum, labium and maxillae as shown in Fig. 217 View Figs 211–219 . Maxillae with 7 cuspules each.

LEGS. Tibia and metatarsus I as in Fig. 274 View Figs 274–281 . Scopula: entire and distal on metatarsi I–II; entire on tarsus I; narrowly divided with setae on tarsus II; widely divided on tarsus III; rudimentary on tarsus IV. Trichobothria: 2 rows of 7 each on tibiae, 10–12 on metatarsi, 7–10 on tarsi, 6 on cymbium. PTC I–IV with 6–7 and 5 teeth on outer and inner margins, respectively.

SPINATION. Palp: femur d3, pd2; tibia d2, p3, r1, v4; cymbium d4. Leg I: femur d4, pd3, rd3; patella p1; tibia p2, pv2, rv2+2M; metatarsus v1. Leg II: femur d4, pd3; patella p1; tibia p3, v6; metatarsus v6. Leg III: femur d4, pd3, rd3; patella p1, r1; tibia d2, p2, r2, v6; metatarsus d1, p3, r3, v7. Leg IV: femur d4, pd3, rd3; patella p1; tibia d2, p3, r3, v7; metatarsus d1, p3, r2, v9(8). Palpal patella and tarsi I–IV aspinose.

PALP. Tibia, cymbium and copulatory bulb as shown in Fig. 366 View Figs 364–378 . Embolus with widely tapering basal portion ending with triangular keel, and with twisted apical part (as shown in Figs 432–433 View Figs 429–438 ).

SPINNERETS. See Fig. 591 View Figs 584–592 . PLS: maximal diameter 0.37; length of basal, medial and apical segments 0.60, 0.42, 0.27; total length 1.29; apical segment triangular.

	Femur	Patella	Tibia	Metatarsus	Tarsus	Total
Palp	1.93 (2.68)	1.07 (1.45)	1.43 (1.69)	–	0.65 (1.42)	5.08 (7.24)
Leg I	3.20 (3.23)	1.78 (2.09)	2.43 (2.35)	2.54 (1.77)	1.45 (1.34)	11.40 (10.78)
Leg II	2.98 (2.96)	1.50 (1.89)	2.18 (2.08)	2.14 (1.84)	1.43 (1.36)	10.23 (10.13)
Leg III	2.57 (2.63)	1.43 (1.54)	1.89 (1.78)	2.60 (2.25)	1.46 (1.39)	9.95 (9.59)
Leg IV	3.51 (3.54)	1.48 (1.92)	2.76 (2.66)	3.75 (3.31)	2.07 (1.61)	13.57 (13.04)

Female (paratype)

HABITUS. See Fig. 51. View Figs 45–53

MEASUREMENTS. TBL 15.05, CL 4.22, CW 3.91, LL 0.41, LW 0.87, SL 2.15, SW 2.06.

COLOUR. Mostly as in male, but chelicerae light scarlet red and legs more uniformly coloured, without difference between legs I and II–IV.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 125 View Figs 118–126 . Clypeus and eye group as in Fig. 183 View Figs 172–183 . Eye diameters and interdistances: AME 0.13(0.17), ALE 0.24, PLE 0.15, PME 0.07; AME–AME 0.13(0.09), ALE–AME 0.09(0.07), ALE–PLE 0.09, PLE–PME 0.06, PME–PME 0.40. Cheliceral rastellum absent. Each cheliceral furrow with 10 promarginal teeth and 5–6 mesobasal denticles. Sternum, labium and maxillae as shown in Fig. 244 View Figs 238–246 . Maxillae with 13–14 cuspules each.

LEGS. Scopula: entire and distal on metatarsi I–II; entire on palpal tarsus; narrowly divided on tarsus I; widely divided by setae on tarsus II; absent on metatarsi and tarsi III and IV. Trichobothria: 2 rows of 9–10 each on tibiae, 13–15 on metatarsi, 11–13 on tarsi, 9–10 on palpal tarsus. Palpal claw with 4 promarginal teeth widely separated from each other. PTC I–II and III–IV with 6–7 and 6–8 teeth on each margin, respectively.

SPINATION. Palpal femur and femora I–II with 4 dorsal bristles; femora III–IV with one thin basodorsal spine and 3 dorsal bristles alongside midline; palpal patella, patellae I–II and tarsi I–IV aspinose. Palp: femur pd1; tibia v10(7); tarsus v4(3). Leg I: femur pd1; tibia v5; metatarsus v6. Leg II: femur pd1; tibia p3, v7; metatarsus p1, v7(6). Leg III: femur pd3, rd2; patella p1, r1; tibia d1, p2, r2, v7; metatarsus d2, p3, r3(2), v7. Leg IV: femur rd1; patella r1; tibia d1, p2, r3, v7; metatarsus d1, p3, r3, v10(9).

SPERMATHECAE. Each of paired spermathecae with wide cone-shaped trunk and short fusiform outer branch ( Figs 528–529 View Figs 522–536 ).

SPINNERETS. See Fig. 592 View Figs 584–592 . PMS: absent. PLS: maximal diameter 0.45; length of basal, medial and apical segments 0.84, 0.55, 0.53; total length 1.92; apical segment triangular.

Variation

Carapace length in males (n= 3) varies from 3.30 to 3.95. One of the fragmented paralectotype males possesses 8 (vs 7 in the lectotype) teeth on the cheliceral furrow promargin ( Fig. 199 View Figs 196–201 ). According to Charitonov (1946), within the type series, the number of the maxillary cuspules ranges from 5 to 8 (some type specimens were used by D.E. Charitonov as preparations, and only a few of their parts have survived to date). A weak difference in the structure of the male tibia and metatarsus I is shown in Fig. 275 View Figs 274–281 . Most other features, including a specific shape of the embolic keel, were found to be consistent throughout the available specimens.

Ecology

Raveniola fedotovi inhabits the rocky midland montane slopes between 1100 and 1600 m a.s.l., from the steppe shrubland zone in high foothills ( Fig. 707 View Figs 707–714 ) to the higher-located zone of mosaic riverside woodland dominated by Juglans regia ( Fig. 708 View Figs 707–714 ). The collected conspecific specimens were found in overgrown screes under shrubs and tree canopies hiding deeply inside the rocky layer ( Figs 709–710 View Figs 707–714 ).

Distribution

South Uzbekistan. See Fig. 755 View Figs 751–760 .

ZMMU	Zoological Museum, Moscow Lomonosov State University
PSU	Portland State University, Vertebrate Biology Museum
SMNH	Department of Paleozoology, Swedish Museum of Natural History

Copyright notice

No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.

Taxonomy

Kingdom	Animalia
Phylum	Arthropoda
Class	Arachnida
Order	Araneae
Family	Nemesiidae
Genus	Raveniola