Raveniola mikhailovi Zonstein, 2021

Zonstein, Sergei L., 2024, A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia, European Journal of Taxonomy 967, pp. 1-185 : 68-71

publication ID	https://doi.org/ 10.5852/ejt.2024.967.2699
publication LSID	lsid:zoobank.org:pub:C08B8027-50CC-417E-BCD4-5183B9FF6738
DOI	https://doi.org/10.5281/zenodo.14026346
persistent identifier	https://treatment.plazi.org/id/03A8B723-5569-FFFC-FDD2-E1CCFAB7CA0B
treatment provided by	Plazi
scientific name	Raveniola mikhailovi Zonstein, 2021
status

Treatment

Raveniola mikhailovi Zonstein, 2021 View in CoL

Figs 23–24 View Figs 19–27 , 56–57 View Figs 54–62 , 78–79 View Figs 76–81 , 104 View Figs 100–108 , 130 View Figs 127–135 , 159 View Figs 148–159 , 188–189 View Figs 184–195 , 223 View Figs 220–228 , 249–250 View Figs 247–255 , 282 View Figs 282–289 , 306 View Figs 290–309 , 373 View Figs 364–378 , 445–447 View Figs 439–447 , 538–540 View Figs 537–554 , 605–608 View Figs 602–610 , 720–722, 758

Raveniola mikhailovi Zonstein 2021: 209 View in CoL View Cited Treatment , figs 1–2, 5–6, 13–15 (♂ ♀), except for the mismatched figs 9–10 (♂; see the corresponding notes below).

Raveniola virgata View in CoL – Zonstein 1987: 1018 (part).

Diagnosis

Males of Raveniola mikhailovi differ from the related male congeners by the following characters: from R. nenilini sp. nov. and R. vulpina sp. nov. by a gently twisted (vs slightly arcuate) embolus, and from R. virgata in having a less stout palpal tibia, as well as a thinner tibia and metatarsus I ( Figs 282 View Figs 282–289 , 373 View Figs 364–378 , 445–447 View Figs 439–447 cf. Figs 286–287 View Figs 282–289 , 376 View Figs 364–378 , 448–449 View Figs 448–456 , 459–465 View Figs 457–465 ). Females of R. mikhailovi are distinguishable due to a specific structure of the spermathecae, with relatively short trunks and widely diverging lateral diverticula (vs differently arranged spermathecal structures in other species; see Figs 538–540 View Figs 537–554 cf. Figs 541–543, 547–554 View Figs 537–554 ).

Material examined

Holotype

KYRGYZSTAN • ♂; Chatkal Mts (southern slope), Hoja-Ata Canyon , Karangitun Gorge ; 41°46′ N, 71°56′ E; 1200–1400 m a.s.l.; 2 May 1983; S. Zonstein leg.; SMNH. GoogleMaps

Paratypes (8 ♂♂, 24 ♀♀, 1 ♀ subad.)

KYRGYZSTAN • 3 ♀♀; same collection data as for preceding; SMNH GoogleMaps • 2 ♂♂, 2 ♀♀; same collection data as for preceding, Tumanyak Gorge ; 41°49′ N, 71°56′ E; 1800 m a.s.l.; 5 Jul. 2000; S. Zonstein leg.; SMNH GoogleMaps • 5 ♀♀, 1 ♀ subad.; same collection data as for preceding, Kokkolot Gorge ; 41°47′ N, 71°57′ E; 1600 m a.s.l.; 16 May 1982; S.V. Ovchinnikov leg.; SMNH GoogleMaps • 4 ♀♀; same collection data as for preceding, Kichkil Gorge ; 41°50′ N, 71°57′ E; 1400 m a.s.l.; 9 Jul. 1983; K.G. Mikhailov leg.; ZMMU GoogleMaps • 5 ♂♂, 1 ♀; same collection data as for preceding, vicinity of Sary-Chelek Lake ; 41°52′ N, 71°58′ E; 1900–2000 m a.s.l.; 28 May 1992; S. Zonstein leg.; SMNH GoogleMaps • 1 ♂, 9 ♀♀; Chatkal Mts , Aflatun Canyon , Oyalma (Uyalma) Gorge ; 41°52′ N, 71°51′ E; 1800 m a.s.l.; 29 Jul. 1983; K.G. Mikhailov leg.; ZMMU GoogleMaps .

Additional material (4 ♀♀, 2 ♀♀ subad.)

KYRGYZSTAN • 2 ♀♀ subad.; Chatkal Mts , Sary-Chelek Reserve ; 25 Jul. 1968; V.F. Bakhvalov leg.; SMNH • 4 ♀♀; Chatkal Mts , Chapchama Pass ; 41°32′ N, 70°49′ E; 2850 m a.s.l.; 8 Jul. 1968; V.F. Bakhvalov leg.; SMNH GoogleMaps .

Description

Male (holotype)

HABITUS. See Fig. 23. View Figs 19–27

MEASUREMENTS. TBL 12.30, CL 4.56, CW 4.12, LL 0.39, LW 0.81, SL 2.33, SW 2.16.

COLOUR. Carapace, palps and legs medium yellowish orange; leg I slightly darker than other legs; eye tubercle blackish brown; chelicerae light cherry red; sternum, labium and maxillae light yellowish orange; abdomen greyish brown, with darker brown dorsal chevron-like pattern and a few small brown marks on ventral side; book-lungs and spinnerets pale yellowish brown.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 104 View Figs 100–108 . Clypeus and eye group as in Fig. 159 View Figs 148–159 . Eye diameters and interdistances: AME 0.15(0.22), ALE 0.27, PLE 0.20, PME 0.18; AME–AME 0.12(0.05), ALE–AME 0.06(0.03), ALE–PLE 0.05, PLE–PME 0.02, PME–PME 0.29. Anterior cheliceral edge with unmodified setae; rastellum not developed. Each cheliceral furrow with 9–10 promarginal teeth and 2 mesobasal denticles. MIT indiscernible. Sternum, labium and maxillae as shown in Fig. 223 View Figs 220–228 . Maxillae with 11–15 cuspules each.

LEGS. Tibia and metatarsus I as in Figs 282 View Figs 282–289 , 306 View Figs 290–309 . Scopula: entire and distal on metatarsi I–II; entire on tarsus I; narrowly divided by setae on tarsus II; sparse and very widely divided on tarsi III–IV.

Trichobothria: 2 rows of 8–9 each on tibiae, 12–14 on metatarsi, 11–12 on tarsi, 8 on cymbium. PTC I–II and III–IV with 8–10 and 9–11 teeth on each margin, respectively.

SPINATION. Palp: femur d3, pd2, rd2; patella pd1; tibia d2, p3, r1, v6; cymbium d10(12). Leg I: femur d4, pd3, rd3; patella p1; tibia p2, pv1, r2, rv2+2M; metatarsus v1. Leg II: femur d4, pd3; patella p1; tibia p4(3), v7; metatarsus p2(1), v4(3). Leg III: femur d4, pd3, rd2; patella p3(2), r1; tibia d3, p3, r3, v8; metatarsus p3, r3, v8. Leg IV: femur d4, pd3, rd3; patella p1, r1; tibia d3, p3, r3, v9; metatarsus d3, p4, r4, v8. Tarsi I–IV aspinose.

PALP. Tibia, cymbium and copulatory bulb as shown in Fig. 373 View Figs 364–378 . Embolus long tapering and slightly curved subapically ( Figs 445–447 View Figs 439–447 ).

SPINNERETS. See Figs 605–606 View Figs 602–610 . PMS: length 0.23, diameter 0.12. PLS: maximal diameter 0.42; length of basal, medial and apical segments 0.68, 0.47, 0.38; total length 1.53; apical segment triangular.

	Femur	Patella	Tibia	Metatarsus	Tarsus	Total
Palp	2.61 (3.19)	1.41 (1.78)	1.83 (2.36)	–	0.72 (2.25)	6.57 (9.58)
Leg I	4.13 (4.31)	2.26 (2.77)	3.36 (3.41)	3.27 (2.66)	1.99 (2.04)	15.01 (15.19)
Leg II	3.81 (4.13)	2.03 (2.47)	3.02 (2.88)	3.07 (2.70)	1.94 (2.03)	13.87 (14.21)
Leg III	3.63 (3.46)	1.71 (1.89)	2.58 (2.45)	3.73 (3.02)	2.03 (2.03)	13.68 (12.85)
Leg IV	4.71 (4.62)	2.14 (2.50)	3.67 (3.59)	5.18 (4.66)	2.65 (2.25)	18.35 (17.62)

Female (paratype)

HABITUS. See Fig. 57. View Figs 54–62

MEASUREMENTS. TBL 18.10, CL 6.56, CW 5.54, LL 0.58, LW 1.13, SL 3.35, SW 2.84.

COLOUR. As in male.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 130 View Figs 127–135 . Clypeus and eye group as in Fig. 189 View Figs 184–195 . Eye diameters and interdistances: AME 0.13(0.19), ALE 0.28, PLE 0.20, PME 0.14; AME–AME 0.14(0.08), ALE–AME 0.12(0.09), ALE–PLE 0.07, PLE–PME 0.06, PME–PME 0.40. Cheliceral rastellum absent. Each cheliceral furrow with 9 promarginal teeth and 3 mesobasal denticles. Sternum, labium and maxillae as shown in Fig. 250 View Figs 247–255 . Maxillae with 12–16 cuspules each.

LEGS. Scopula: entire and distal on metatarsi I–II; entire on palpal tarsus and tarsus I; narrowly divided by setae on tarsus II; sparse and widely divided on tarsus III; rudimentary on tarsus IV. Trichobothria: 2 rows of 9–11 each on tibiae, 15–16 on metatarsi, 14–15 on tarsi, 10 on palpal tarsus. Palpal claw with 4 long promarginal teeth. PTC I–II and III–IV with 6–7 and 7–9 teeth on each margin, respectively.

SPINATION. Femora with 1–2 basodorsal spine and 2–3 dorsal bristles; palpal patella and tarsi I–IV aspinose. Palp: femur pd1; tibia v7; tarsus v4. Leg I: femur pd1; patella p1; tibia p2, v7; metatarsus v6. Leg II: femur pd1; patella p1; tibia p2, v7; metatarsus v6. Leg III: femur pd3, rd2; patella p2, r1; tibia d1, p2, r2, v7; metatarsus d1, p4, r3, v8(7). Leg IV: femur pd1, rd1; patella p1, r1; tibia d1, p3(2), r3, v7; metatarsus d1, p4, r4, v12(10).

SPERMATHECAE. Each of paired spermathecae Y-shaped with relatively short and wide base carrying two relatively short and widely diverging branches ( Fig. 540 View Figs 537–554 ).

SPINNERETS. See Fig. 608 View Figs 602–610 . PMS: length 0.38, diameter 0.18. PLS: maximal diameter 0.62; length of basal, medial and apical segments 1.08, 0.55, 0.48; total length 2.11; apical segment triangular.

Variation

Carapace length in males (n=9) varies from 4.41 to 5.69, in females (n=11) from 4.72 to 7.37. Variations in the habitus, the eye group arrangement, and the structure of the sternum and the spinnerets are shown in Figs 24 View Figs 19–27 , 56 View Figs 54–62 , 78–79 View Figs 76–81 , 188 View Figs 184–195 , 249 View Figs 247–255 , 607 View Figs 602–610 . Variation in the structure of the spermathecae as shown in Figs 538–540 View Figs 537–554 .

Ecology

According to the observations and the labelled collection data, the spiders were collected under stones in a wide array of montane habitats – from shrubland on the lower border of the forested zone via the broad-leaved, mixed and coniferous montane forests (dominated by Juglans regia and Picea schrenkiana Fisch. & C.A. Mey. , respectively) to the subalpine and alpine woodless grasslands ( Figs 720–722 View Figs 715–722 ). The spiders use cavities under stones as retreats. In Sary-Chelek Reserve, they can occur together with a sympatric species, Raveniola vulpina sp. nov. (the ranges of these two species partially overlap).

Distribution

Known from Western Tien-Shan: Chatkal Mts. See Fig. 758 View Figs 751–760 .

Notes

When describing this species, the illustrations showing the copulatory bulb of the male holotype of R. vulpina sp. nov., stored in a folder under the name indicating the type locality, common for two species (Sary-Chelek), were mistakenly used instead of images of this structure actually belonging to the male holotype of the sympatric R. mikhailovi , kept in the same folder. This error is corrected herein. All other images of the holotype, used at the original description ( Zonstein 2021: figs 1, 5), are correct.