Chiridopsis aubei ( Boheman, 1855 )

Świ, Jolanta, 2017, A monograph of the Afrotropical Cassidinae (Coleoptera: Chrysomelidae). Part 4. Revision of the genus Chiridopsis Spaeth, Zootaxa 4316 (1), pp. 1-85 : 12-22

Chiridopsis aubei ( Boheman, 1855)

( figs 2 , 21 , 31–42 )

Coptocycla Aubei Boheman, 1855: 195, 1857: 169 , 1862: 419; Gemminger and Harold, 1876: 3666; Węgrzynowicz and Wąsowska, 1996: 43.

Chirida aubei: Weise, 1896 a: 9 ; 1896 c: 23, 24; Spaeth, 1903: 179, 1914: 126, 1916: 41.

Chiridopsis aubei: Spaeth, 1924: 349, 1929: 161 , 1935: 175, 1943: 62; Shaw, 1956: 596, 1963: 460, 1968 a: 371, 1968 b: 783, 1972: 78; Borowiec, 1985 a: 241; 1985 b: 448, 1986: 807, 1999: 297; Nummelin and Borowiec, 1991: 14.

Chiridopsis aubei ab. dissoluta Spaeth, 1933: 357, 1935: 175; Shaw, 1961: 34, infrasubspecific unavailable name.

Coptocycla semivittata Boheman, 1855: 194, 1857: 168 , 1862: 418; Gemminger and Harold, 1876: 3674; Węgrzynowicz and Wąsowska, 1996: 44; Borowiec, 1999: 297 (as syn. of Ch. aubei ).

Chirida semivittata: Weise, 1896 c: 23 , 24; Spaeth, 1914: 127.

Coptocycla mesomela Boheman, 1862: 426 ; Borowiec, 1999: 297 (as syn. of Ch. aubei ).

Coptocycla mesomelaena [sic]: Gemminger and Harold, 1876: 3671; Spaeth, 1914: 131.

Chiridopsis mesomelaena [sic]: Spaeth, 1924: 347.

Coptocycla fausta Boheman, 1862: 420 ; Gemminger and Harold, 1876: 3669; Spaeth, 1914: 126 (as syn. of Ch. aubei ).

Chirida fausta: Weise, 1896 c: 23 , 24 (as syn. of Ch. aubei ).

Chirida Baumanni Spaeth, 1902: 459, 1909: 276 , 1914: 126; Borowiec, 1986: 807 (as syn. of Ch. aubei ).

Chiridopsis baumanni: Spaeth, 1924: 338 , 342, 1925: 7, 1943: 62; Shaw, 1961: 34, 1968 b: 783, 1972: 78.

Chiridopsis baumani [sic]: Shaw, 1955: 238, 1963: 461.

Chiridopsis Baumanni ssp. nigriceps Spaeth, 1924: 343 ; Shaw, 1955: 238 (as possible syn. of Ch. baumanni ); Borowiec, 1986: 807 (as syn. of Ch. aubei ).

Chirida breviuscula Weise, 1904: 55 ; Spaeth, 1914: 126; Borowiec, 1999: 297 (as syn. of Ch. aubei ).

Chiridopsis breviuscula: Spaeth, 1924: 349 ; Shaw, 1972: 78 (incl. fig.).

Chiridopsis signaticollis Spaeth, 1924: 346 ; Borowiec, 1986: 807 (as syn. of Ch. aubei ).

Chiridopsis aubei ab. signaticollis: Spaeth, 1933: 357 .

Chiridopsis aubei var. signaticollis: Shaw, 1955: 239 .

Chiridopsis maculipennis Spaeth, 1924: 347 ; Borowiec, 1999: 297 (as syn. of Ch. aubei ).

Chirida Gyldenstolpei Weise, 1924: 27 ; Borowiec, 1999: 297 (as syn. of Ch. aubei ).

Chiridopsis Gyldenstolpei : Spaeth, 1925: 6.

Chiridopsis Weisei Spaeth, 1924: 349 (new substitute name for C. semivittata sensu Weise, 1896 not Boheman, 1855); Borowiec, 1999: 305, n. syn.

Description. Le: 4.4–6.25 mm, Wi: 3.4–4.8 mm, Lp: 1.6–2.25 mm, Wp: 2.7–3.55 mm, Le/Wi: 1.20–1.35, Wp/Lp ratio: 1.50–1.66. Body from short–oval to oval, males slightly stouter than females.

In typically coloured specimens pronotal disc black with two smal yellow spots at base, scutellum black, elytral disc yellow with three broad, black vittae, along suture and along sides, connected on slope but marginal interval and apex of disc yellow ( figs 31–33 ). In common form named ab. baumanni dorsum uniformly yellow ( fig. 39 ). Between these forms many intermediates were observed ( figs 34–38 ): often black on pronotal disc forms only triangular spot with two large basal yellow spots ( figs 34, 36 , 37 ) or is reduced to three basal stripes ( fig. 35 ), elytral vittae sometimes narrow and not connected apically ( figs 34–36 ), or reduced to short rudiments of lateral vittae or spots at base and posterolateral parts of disc ( fig. 37 ), occasionally only pronotum with M-shaped pattern but elytra uniformly yellow ( fig. 38 ). In southern parts of distribution range predominate forms with pronotal pattern reduced to three or four spots and elytral lateral vittae divided into 3–5 separate spots ( figs 40–42 ), sometimes also separate spot on humeral callus and small additional spot in 1/5 length of second interval ( fig. 40 ), sutural vitta from broad with irregular sides to very narrow, limited to sutural margin, or completely obsolete (= ab. breviuscula ). Explanate margin always yellow. Head usually black, sometimes brown, or area between clypeal grooves brown and central plate black, thorax black, abdomen black narrowly surrounded by yellow. Coxa usually black, trochanters more or less infuscate, rest of legs yellow. Antennae yellow, sometimes last segment infuscate apically.

Pronotum elliptical ( figs 31, 32, 34–42 ), with maximum width in the middle, anterior margin regularly convex, sides broadly rounded, no basal corners. Surface of disc impunctate or with extremely fine sparse pricks, shiny. Explanate margin broad, impunctate, shiny, transparent with well visible honeycomb structure.

Base of elytra much wider than base of pronotum ( figs 31, 32, 34–42 ), humeral angles distinctly protruding anterad, subangulate, in some specimens elytral margin behind humeral angle shallowly emarginated, then humeri subacute. Disc regularly convex in profile with top of convexity in postscutellar point ( fig. 33 ), without impressions. Punctation fine, arranged in regular rows, from sparse to moderately dense, distance between punctures from as wide as to three times wider than puncture diameter. Marginal row distinct, its punctures distinctly coarser than punctures in central rows. Intervals flat, three to five times wider than rows, marginal interval 1.5–2.0 times wider than submarginal one, no humeral or lateral folds. Explanate margin narrow, strongly declivous, in the widest part 5.5–6.0 times narrower than disc, surface usually impunctate but in some specimens with very shallow sparse punctures, shiny, transparent with well visible honeycomb structure.

Eyes very large, gena obsolete ( fig. 21 ). Clypeus broad, approximately 1.3 times as wide as long. Clypeal grooves fine but well marked, in most populations in basal 2/3 length of clypeus running close to margin of eye then converging in obtuse angle or arch, in populations from extreme west and extreme east of range clypeal grooves run close to margin of eye only in basal half then converging in obtuse angle, surface of clypeus from slightly to strongly convex, the most convex clypeus has population from Cameroon and bassin of Congo, the less convex populations from extreme east. Surface of clypeus microreticulate, from slightly alutaceous to slightly shiny with few very small, setose punctures. Labrum minutely emarginate to 1/6 length. Antennae stout, segments 9–10 slightly transverse. Length ratio of antennal segments: 100:59:76:76:65:59:56:53:56:59:103. Segment 3 approximately 1.3 times as long as segment 2 and as long as segment 4.

Prosternum moderately broad in the middle, moderately expanded apically, area between coxa flat but with more or less developed median groove, surface from regular to slightly obliquely wrinkled on sides with several small hairy punctures, expanded apex not or slightly convex with slightly irregular surface and few coarse, hairy punctures.

Claws with small basal tooth.

Distribution. Widespread species, known from West, Central and East Africa, south to Zambia ( fig. 2 ). In forest regions of western and central Africa predominates stripped nominotypical form ( figs 31–33 ), in arid sub- Saharan Region and eastern Africa predominates unicolorous form ab. baumanni ( fig. 39 ).

Remarks. One of the most widely distributed and common African cassids, very variable in body shape and colour pattern. Generally, in Central Africa in wet habitats predominates vittate form ( figs 31–33 ), while in sub- Saharan Region and eastern Africa in dry habitats predominates uniformly yellow form ( fig. 39 ) but sometimes within vittate populations occur specimens of unicolorous form. Specimens from western part of range are usually stouter, with more acute humeri than specimens from eastern part of range. Clypeus in western populations is much more convex than in eastern forms, although eastern populations have much more impressed clypeal lines. The relation among various forms of Ch. aubei needs genetic studies in the field and in laboratory conditions. Perhaps the taxon is a group of sibling species with partly overlapping characters, or its polymorphism has a partly geographic character. Dried museum specimens do not allow to verify this hypothesis. Thus, we treated all forms of Ch. aubei -complex as a single polymorphic species. On the other hand, several uniformly yellow taxa described from eastern and southern part of central Africa are very close to particular populations of Ch. aubei and their specific status is also doubtful.We have conserved species status for most of them because examined material of these taxa is not as rich in specimens as Ch. aubei . Their relationships to Ch. aubei also need genetic studies.

Spaeth (1924) suggested that Ch. semivittata sensu Weise, 1896 is not conspecific with C. semivittata Boheman, 1855 , and proposed for it a new substitute name Ch. weisei Spaeth, 1924 . We have examined specimens preserved in ZMHU and identified by Weise as Ch. semivittata and in our opinion they really belong to a different aberration than specimens named as semivittata by Boheman (1855). On the other hand, both represent variations of the extremely variable Ch. aubei (Boh.) . Thus, the name Ch. weisei is a junior synonym of Ch. aubei . Maculate form was described as Ch. aubei ab. dissoluta ( figs 40–42 ) but vittate, maculate and uniformly yellow forms often occur simultaneously in one locality thus we treated Ch. aubei ab. dissoluta as simple synonym of Ch. aubei . Maculate forms predominate in southern part of range of this wide-spread and variable taxon.

The typical vittate form of Ch. aubei ( figs 31, 32 ) is very similar to untypical form of Ch. quadriguttata ( figs 105, 106 ) with regular elytral stripes, but in Ch. aubei inner margin of lateral stripes is always regular while in Ch. quadriguttata the margin in mid length is at least slightly bent towards the suture. Form of Ch. aubei with thin lateral stripes not connected on suture ( figs 34, 35 ) is similar to Ch. circe ( fig. 48 ) but differs in fine elytral punctation with interspaces in lateral rows mostly twice as wide as puncture diameter while in Ch. circe punctation of elytra is coarse and dense with punctures in lateral rows almost touching each other. Small specimens of Ch. aequinoctialis are similar to the vittate form of Ch. aubei ( figs 24, 25 , 31, 32 ) but differ in yellow clypeus and black marginal interval while in Ch. aubei clypeus is black and marginal interval yellow ( figs 19 , 21 ). Spotted form of Ch. aubei (described as ab. dissoluta) is similar to the group of species with pattern formed by rows of separate black spots: Ch. nigrostillata , Ch. tanganikana and Ch. tessellata ( figs 40–42 , 83, 84 , 96–101 ) but they differ in presence of at least two pairs of black spots behind scutellum while in Ch. aubei no spot occurs in sutural area; at most along the suture runs a more or less regular black vitta. The most similar pattern to spotted form of Ch. aubei has Ch. picturata ( figs 94, 95 ) but it differs in more elongate body (length/width ratio 1.48–1.49 vs. 1.29–1.32), base of elytra only slightly wider than pronotum and strongly declivous explanate margin of elytra. Completely yellow form of Ch. aubei (described as Ch. baumanni ) is very similar to two other yellow species with black clypeus and short-oval to subcircular body: Ch. observabilis and Ch. flavipennis ( figs 39 , 56–61 , 111, 112 ). Identification of these three forms needs comparison with series of properly identified specimens. In Ch. aubei clypeal grooves run close to margin to eyes and on top of clypeal triangle converging in obtuse angle while in two relatives clypeal grooves form regular triangle with sharp top thus between the triangle and inner margin of eyes runs well visible, convex fold often paler coloured than the clypeal triangle. Of these three similar forms Ch. aubei ab. baumanni has the most convex clypeus, especially in western populations the convexity is very high while in both relatives the clypeus is flat or only moderately convex.

Types examined. Holotype of Cassida aubei : “Celebes [sic!]” “corr. Calabar” (NRS).

2 syntypes of Coptocycla semivittata : [ NIGERIA] Old Calabar (1 BMNH, 1 NRS).

3 syntypes of Chiridopsis signaticollis : [ IVORY COAST:] Aknopong (MM).

Holotype of Chiridopsis maculipennis : [ ZAMBIA] NW Rhodesia (MM).

Syntype of Chirida baumanni : DEMOCRATIC REPUBLIC OF CONGO ( ZAIRE) Mpala (MM)

1 syntype of Chiridopsis baumanni ssp. nigriceps : [UGANDA] Tero Forest, VII 1912, C.C. Gowdey (BMNH); 19 syntypes of Chiridopsis baumanni ssp. nigriceps : [CAMEROON] Ekok, A. Heyne (MM).

Holotype of Chirida breviuscula : [ KENYA] Ikutha ( ZMHU).

1 syntype of Chirida gyldenstolpei : DEMOCRATIC REPUBLIC OF CONGO (ZAIRE) Ituru, Guld. Exp. (NRS); 1 syntype of Chirida gyldenstolpei : DEMOCRATIC REPUBLIC OF CONGO (ZAIRE) Ituru, Conradt (MM).

1 syntype of Chiridopsis aubei ab. dissoluta: DEMOCRATIC REPUBLIC OF CONGO ( ZAIRE) Kabambare , 1 III 1907, H. Wilmin ( MRAC) ; 2 syntypes of Chiridopsis aubei ab. dissoluta: DEMOCRATIC REPUBLIC OF CONGO ( ZAIRE) Kasongo, Niembo , VI 1918, R. Mayné ( MRAC) ; 1 syntype of Chiridopsis aubei ab. dissoluta: DEMOCRATIC REPUBLIC OF CONGO ( ZAIRE) Kikwit , 1920, P. Vanderijst ( MRAC) ; 3 syntypes of Chiridopsis aubei ab. dissoluta: DEMOCRATIC REPUBLIC OF CONGO ( ZAIRE) Kisantu, Vanderijst (2 MRAC, 1 MM) ; 1 syntype of Chiridopsis aubei ab. dissoluta: DEMOCRATIC REPUBLIC OF CONGO ( ZAIRE) Kisantu, 1919, Vanderijst ( MRAC) ; 1 syntype of Chiridopsis aubei ab. dissoluta: DEMOCRATIC REPUBLIC OF CONGO ( ZAIRE) Lemfu, Vanderijst ( MRAC) ; 1 syntype of Chiridopsis aubei ab. dissoluta: DEMOCRATIC REPUBLIC OF CONGO ( ZAIRE) Lemfu à Kampese, P. Vanderijst (MM) ; 3 syntypes of Chiridopsis aubei ab. dissoluta: DEMOCRATIC REPUBLIC OF CONGO ( ZAIRE) V 1918, R. Mayné ( MRAC) ; 9 syntypes of Chiridopsis aubei ab. dissoluta: DEMOCRATIC REPUBLIC OF CONGO ( ZAIRE) Stanleyville , X 1925, J. Ghesquière ( MRAC) .

Coptocycla mesomela Boheman, 1862 and Coptocycla fausta Boheman, 1862 were desribed from “Calabar” (now Nigeria) but their type location is unknown.

Other specimens examined. ANGOLA: Kabinda, 1935, 1 ab. dissoluta, P. Henrard ( MRAC) ; Huila, Tschivinguire, 14–17 XI 1974, 1 ab. baumanni (WM) ; Luanda, 1965, 6 + 1 ab. baumanni ), Giraudet ( MRAC); Lunda, 1200 m, 1927, 1 ab. dissoluta, P. Nannings (ITZ); Quitondo, Distr. Calulo, 11 VIII–2 IX 1967, 2 + 2 ab. dissoluta (ACO).

BENIN: Cove , Zou River env., 16.IV.2000, 2, Z. Andrs (FK); Dahomey, Zagnanano, 1, Le Moult ( IRSN).

BURKINA FASO: Haute Volta, Melou , 7 ( IRSN).

BURUNDI: Bugesera, rives lac Tshohoha , IX 1957, 1 ab. baumanni, N. Leleup ( MRAC) ; Bururi, 1 ab. baumanni, Giraudin ( MRAC) ; Cibitoke, II 1989, 3, C.J.M. Berger (RB); Kibira-Pare, Teza, 29 II 1992, 1, Arndt ( NME) ; Plaine de la Ruzizi , IV 1966, 1, N'Dani ( MRAC) .

CAMEROON: Abong M'Bang, 1–30 IV 1936, 1, F. G. Merfield (BMNH), 1959, 1, R. Lenczy (HNHM); Bamum, VI 1911, 1 (HNHM); Bare, 800 m, 6 XI 1912, 4, Rothkirch (ZMHU); Bare-Dschang, 2–6 XII 1908, 1, Riggenbach (ZMHU); Barombi, 1, Zeuner (ZMHU); Barombi-Likoko road, 20 I 1958, 1, W. Hartwig (MKB); Batanga, 11 I 1911, 1, J.A. Reis (CMNH), I 1911, 2, 12 V 1911, 1, V 1911, 1, VI 1911, 2, A.I. Good (CMNH), IV 1914, 1, V 1914, 1, F.H. Hope (CMNH); Batouri Distr., 750 m, 1 V–6 VI 1935, 3 (BMNH); Batouri, 3.75°N, 13.75°E, 1 V–6 VI 1935, 40, F. G. Merfield (BMNH, LS); Batouri, Savanah Country, 1 I – 24 II 1936, 1, F. G. Merfield (BMNH); Bipindi, 20, Zenker (ZMHU); Bubandjidole, Tibati, 15 VI–7 VII 1901, 1, Glauning (ZMHU); Buea, 1–7 III 1912, 1, Rothkirch (ZMHU); Dimako-Abong Mbang, 13 VI 1973, 1, R. Linnavuori (ZMUH); D'Ja Posten, 3.15°N, 13.30°E, 15 V–1 VII 1936, 2, F. G. Merfield (BMNH); Djoungolo, Nkongzok, 11 II 1963, 2, L. Segers (ZSM); Djoungolo, Ntouessong-r. Nsolé, 17 IX 1963, 3 ab. baumanni, L. Segers (ZSM); Djoungolo Arr., Villa Carde n. Nyong, 23 VII 1963, 2, 28 VIII 1963, 1 ab. baumanni (ZSM); Douala, 4, Lencsz, 1, Cantaloube (MRAC); Doumé, II 1959, 1 ab. baumanni (MRAC); Duala, IX 1912, 2, Rothkirch (ZMHU); Ebolowe, 700 m, 15– 25 IV 1912, Rothkirch (ZMHU); Edea, 15 III 1 922, 1, 25 III 1922, 1, 9 V 1922, 1, 9 IX 1922, 1, 25 IX 1923, 1, J.A. Reis (CMNH), 2, Carret (MRAC); Ediki, 21 VI 1973, 1, R. Linnavuori (ZMUH); Efulen, IX 1912, 4, XI 1912, 12, J.A. Reis (CMNH), 12 IX 1911, 1, 13 IX 1911, 1, 13 I 1918, 1, 17 I 1918, 1, 19 I 1918, 2, H.L. Weber (CMNH); Ekabita, 25–29 IV 1977, 1, Benjamin (HPA); Ekododo, 26 VIII–16 IX 1913, 1, Escherich (ZMHU); Jaunde, 19–23 X 1914, 2 ab. baumanni , Teßmann (ZMHU), 20 III 1 923, 3, 27 III 1923, 2, 1 V 1923, 1 (CMNH); Jaunde- Akonolinga, km 90–94, 2, Naumann (ZMHU); Joh.-Albrechtshöhe, 1 IX 1897 –22 XI 1898, 10 (ZMHU); Joko, 4 (ZMHU), 1 ab. baumanni (HNHM); Kamerunberg, Soppo, 730 m, 1912, 1, v. Rothkirch (FMNH); Katho-Barombi See, I 1958, 2, Hartwig (MKB); Kribi, 5 + 3 ab. baumanni, Carret (MRAC); Kribi, 0–30 m, 18–31 III 1974, 1, M. Kühbaudner (ZSM), 19 VI 2002, 1, Bačovský (LS); Kumbu-Mamfe, 23 VI 1973, 1, R. Linnavuori (ZMUH); Lolodorf, 1, Heyne (ZMHU), V 1914, 2, 8 VI 1914, 2, J.A. Reis (CMNH), 20 V 1914, 1, X 1914, 1, I 1925, 1, III 1925, 3, IV 1925, 1, V 1925, 1, VI 1925, 9, X 1926, 2, XI 1926, 18, A.I. Good (CMNH); Malange, 1, Mechow (ZMHU); Malende, Mt. Kamerun, XII 1957, 3, W. Hartwig (MKB); Malange, 1, Mechow (ZMHU), 1, Pogge (ZMHU); Mbalmayo, 19 VI 1976, 1, C. Besnard (BB); Centre prov., road to Mbankomo, 8 km SW Yaounde, 2400–2600 ft., 24 X 2000, 1, W.J. Bicha (YM); Mfok, Nkolusala, r. Nsala, 27 VIII 1963, 2, L. Segers (ZSM); Mokundange, 1–15 VI 1905, 2, G. Tessmann (ZMHU); Mt. Balmayo, 6, Barga, XII 1962, 1 ab. baumanni, Barga , 1, Cantaloube (MRAC); Mt. Kamerun, 4 (ZMHU); Mueli, Mt. Kamerun, 600 m, II 1958, 5, Hartwig (MKB); Mundame, 1, R. Rohde (ZMHU); Namiong n. Lolodorf, 1, O. Ulrich (ZMHU); Nanga Eboko, VII–X 1959, 1 ab. baumanni, R. Lenczy (HNHM); Nkolbisson, Dept. Nyong-Sanaga, IX 1963, 2, X 1963, 4 + 1 ab. baumanni, L. G. Segers (MRAC); Nkolbison, Yaounde-Bi, 16 II 1963, 2, 20 V 1963, 3 + 1 ab. baumanni, L. Segers (ZSM); N'Kongsamba, 2, J. Cantaloube (MRAC); Nssanakang, 2, A. Dichl (ZMHU); Nulang, VI 1911, 9, L. Colin (ZMHU); Obala, Nkometon, r. Nkoa, 22 VIII 1963, 4, L. Segers (ZSM); Okola, 29 VII 1968, 1, B. de Miré (HPA); Okola, Ebongsi-r. Mbanize, 19 VII 1963, 1 + 1 ab. baumanni , 17 IX 1963, 1, L. Segers (ZSM); Okola, Etoud r. Tchazoumon, 16 IX 1963, 1, L. Segers (ZSM); Okola, Mbanize, 19 VII 1963, 2, L. Segers (ZSM); Okola, Nkollekie-r. Lékie, 14 VIII 1963, 1, L. Segers (ZSM); Okola, Nkong-r. Yégué, 14 VIII 1963, 2 + 2 ab. baumanni, L. Segers (ZSM); Pipinde, 3 for. typ., Zenker (ZMHU); Rei, Bubaudjidda, Tibati, 15 VI–7 VII 1901, 1 ab. baumanni, Glauning (ZMHU); Soppo, 25 II 1912, 2, II 1913, 1, Rothkirch (ZMHU); Sud prov., Bigpaga, 12.5 mil. NE Kribi, 50–150 ft., 29 X 2000, 2, W.J. Bicha (YM); Sud-Ouest Prov., 3 mls. NW Ekona, 1400–2230 ft., 4 XI 2000, 1, W.G. Bicha (YM); Umbal, VI 1905, 1, Glauning (ZMHU); Victoria, 1 (ZMHU); 10 km NW of Victoria, 22 I 1978, 2, Lund University Exp. (LU); Yaounde, X 1923, 9 (IRSN); Yaunde, 27 III 1923, 3 ab. baumanni , 4 IV 1923, 1 ab. baumanni , 9 IV 1923, 1 ab. baumanni , 1 V 1923, 1 ab. baumanni (CMNH); 120 km NW Yaounde, Rt. n. 4, 5 XI 1987, 1, F. Kaplan (TAU).

DEMOCRATIC REPUBLIC OF CONGO (ZAIRE): Albert Nat. Park, Bitshumbi, 10–14 I 1936, 1, H. Damas (MRAC); Albert Nat. Park, Mont Hoyo, riv. Issehe, affl. Bombuo, 1200 m, 18 VII 1955, 1, 26 VII 1955, 1, P. Vanschuytbroeck (MRAC); Albert Nat. Park, Kamande, 926 m, 18 IX 1935, 3 ab. baumanni, H. Damas (MRAC); Albert Nat. Park, Massif Ruwenzori, Bomboka près Kyandolire, 1650 m, 15 X 1952, 1 ab. baumanni , 22 X 1952, 1 ab. baumanni, P. Vanschuytbroeck and J. Kekenbosch (MRAC); Albert Nat. Park, Massif Ruwenzori, Kalonge, riv. Nyamwamba, affl. dr. Butahu, 2150 m, 18 VI 1957, 1, P. Vanschuytbroeck (MRAC); Albert Nat. Park, Mont Hoyo, riv. Issehe, affl. Bombuo, 1200 m, 18 VII 1955, 2 ab. baumanni (MRAC); Albert Nat. Park, Mont Hoyo, grotte Saga-Saga, 1160 m, 3 VIII 1955, 1, P. Vanschuytbroeck (MRAC); Albert Nat. Park, Mont Hoyo, grotte Yolohafiri, 1030 m, 12–17 VII 1955, 1 + 1 ab. baumanni , 19–20 VII 1955, 6 (1 ab. baumanni ), 25–28 VII 1955, 3, 8–9 VIII 1955, 1, P. Vanschuytbroeck (MRAC); Albert Nat. Park, Mutsora, 1939, 7 + 12 ab. baumanni, Hackars (MRAC); Albert Nat. Park, Plaine Semliki, 900–1100 m, IV–X 1937, 1 ab. baumanni, Hackars (MRAC); Albert Nat. Park, Secteur Nord, riv. Butahu, affl. Semliki, 1325 m, 25 VI 1957, 1 ab. baumanni, P. Vanschuytbroeck (MRAC); Albert Nat. Park, Secteur Nord, Mantiba,, Moyenne-Lume, 1260 m, 12 XII 1956, 1, P. Vanschuytbroeck (MRAC); Albert Nat. Park, Secteur Nord, riv. May ya Moto, affl. g. Talya, 1180 m, 9 V 1957, 2 ab. baumanni, P. Vanschuytbroeck (MRAC); Albert Nat. Park, Secteur Nord, Mulingo, Sec. Kikura, Rég. Baniangala, 1350 m, 9 VII 1954, 1 + 3 ab. baumanni , 16 VII 1954, 1 ab. baumanni, P. Vanschuytbroeck and H. Synave (MRAC); Albert Nat. Park, Secteur Nord, riv. Ngokoi, affl. Talya, 1100 m, 27 V 1957, 1 ab. baumanni, P. Vanschuytbroeck (MRAC); Albert Nat. Park, Secteur Tshiaberimu, r. Busiangwa, affl. Musavaki, 2380 m, 11–23 III 1954, 1, P. Vanschuytbroeck and H. Synave (MRAC); Albert Nat. Park, Secteur Tshiaberimu, Hintumo, 2450 m, 24 III 1954, 2 ab. baumanni, P. Vanschuytbroeck and H. Synave (MRAC); Albert Nat. Park, Secteur Tshiaberimu, riv. Kalivina, affl. Talia Nord, 2350 m, 27 VIII 1953, 1 ab. baumanni, P. Vanschuytbroeck and V. Hendrickx (MRAC), 28 III 1 954, 1 ab. baumanni , 29 III 1954, 5 ab. baumanni, P. Vanschuytbroeck and H. Synave (MRAC); Albert Nat. Park, Secteur Tshiaberimu, Kirungu, 2720 m, 11 III 1954, 1 ab. baumanni , 24–26 III 1954, 1 ab. baumanni, P. Vanschuytbroeck and H. Synave (MRAC); Albert Nat. Park, Secteur Tshiaberimu, riv. Kisesa, affl. Talya Nord, 2580 m, 20 IV 1955, 1 ab. baumanni, P. Vanschuytbroeck and R. Fonteyn (MRAC); Albert Nat. Park, Secteur Tshiaberimu, Mbolu- Musavaki, 2150 m, 1 ab. baumanni , 15 IV 1955, 1 ab. baumanni, P. Jolivet (MRAC); Albert Nat. Park, Secteur Tshiaberimu, riv. Mbulikerere, affl. Kalivina, 2720 m, 19 IV 1955, 1 ab. baumanni, P. Jolivet (MRAC); Albert Nat. Park, Secteur Tshiaberimu, riv. Mbulikerere, affl. dr. Talia Nord, 2720 m, 26 VIII 1 953, 1 ab. baumanni, P. Vanschuytbroeck and V. Hendrickx (MRAC); Albert Nat. Park, Secteur Tshiaberimu, Mont Buliwa, 2450 m, 25 IV 1955, 1, P. Vanschuytbroeck and R. Fonteyn (MRAC); Albert Nat. Park, Secteur Tshiaberimu, Mont Musienene, près de Kirungu, 2680 m, 21 IV 1955, 2, P. Vanschuytbroeck and R. Fonteyn (MRAC); Albert Nat. Park, Secteur Tshiaberimu, Mont Musimba, riv. Musavaki, 2450 m, 18 IV 1955, 1 ab. baumanni, P. Vanschuytbroeck and R. Fonteyn (MRAC); Albert Nat. Park, Secteur Tshiaberimu, riv. Musavero, affl. Kitevia, 2200 m, 25 IV 1955, 1, ab. baumanni, P. Jolivet (MRAC); Albert Nat. Park, Secteur Tshiaberimu, Sommet du Mont Tshiaberimu, 3000 m, 22– 24 VIII 1953, 1 ab. baumanni, P. Vanschuytbroeck and V. Hendrickx (MRAC); Albert Nat. Park, Secteur Tshiaberimu, riv. Talia Nord, 2340 m, 23 III 1954, 3 ab. baumanni, P. Vanschuytbroeck and H. Synave (MRAC); Albert Nat. Park, Secteur Tshiaberimu, Vusiango, 9 III 1954, 3 ab. baumanni, P. Vanschuytbroeck and H. Synave (MRAC); Albert Nat. Park, SL Eduard, Kamande, 925 m, 1 X 1935, 1 ab. baumanni , 8 IV 1936, 2 ab. baumanni, L. Lippens (MRAC); Albert Nat. Park, SL Eduard, Rwindi, 25 IV 1936, 1, ab. baumanni, L. Lippens (MRAC); Albertville, XII 1918, 1, R. Mayné (MRAC); Arebi, Bondo Mabé, 24 VII 1925, 2, H. Schouteden (MRAC); Aruwimi, I 1926, 3, J. Ghesquière (MRAC); Bakala-Wamba, 1925, 1 ab. dissoluta, P. Vanderijst (MRAC); Bakusu, IX 1912, 13, R. Mayné (MRAC); Bamania, 1936, 1, P. Longinus (MRAC); Bambesa, 10 IX 1933, 2, 20 IX 1933, 2, J. Leroy (MRAC), I 1934, 2, H.J. Bredo (MRAC), 16 III 1 937, 4, 24 III 1933, 2, 7 IV 1937, 1, 19 IV 1937, 1, 24 IV 1937, 1, 6 V 1937, 1, 10 V 1937, 1, 11 V 1937, 3, 31 V 1937, 2, 15 VI 1937, 1, 21 VI 1937, 1, 23 VI 1937, 2, 24 VI 1937, 1 + 1 ab. baumanni , 29 VI 1937, 1 + 1 ab. baumanni , VII 1937, 1, 14 VII 1937, 3, 28 VII 1937, 1, VIII 1937, 2, 1 IX 1937, 1, X 1937, 2, 30 X 1937, 1, XI 1937, 3, II–III 1938, 1, 16 II 1938, 13, IV 1938, 1, X 1938, 3, 4 X 1938, 1, 5 X 1938, 1, 4 III 1939, 1, 6 III 1 939, 1, IV 1939, 4, 4 IV 1939, 2, 14 V 1939, 2, 7 VI 1939, 1, 11 XI 1939, 1, 18 XII 1939, 1 + 1 ab. baumanni , 20 XII 1939, 1, 22 XII 1939, 3, 26 XII 1939, 2, 28 XII 1939, 5, 3 I 1940, 2, 4 I 1940, 1, 6 I 1940, 2, 8 I 1940, 1, J.M. Vrijdagh (IRSN, MRAC, DBET); Bangala, Bunga, 8 III 1932, 2, H.J. Brédo (MRAC); Barumbu, VII 1925, 2, J. Ghesquière (MRAC); Bas Congo: Mayidi, 1942, 1, Van Eyen (MRAC); Basongo, X 1921, 7, L. Achten (MRAC); Bas Uele, Beo, 20 IV 1958, 2 (FMNH); Bas Uele, Buta, 1926, 1, Fr. Joseph (MRAC); Bas Uele, Djamba, 17–25 XII 1924, 1, H. Schouteden (MRAC); Bas Uele, Koteli, 1–21 I 1929, 1, H. Schouteden (MRAC); Beni, 18 VII 1932, 5, H.J. Brédo (MRAC); Beni n. Lesse, VII 1911, 3, Murtula (MRAC); Bikoro, 2 III 1921, 1, H. Schouteden (MRAC); Binga, 5–12 III 1932, 9, H.J. Brédo (MRAC); Biruwe a Matenda, 16 IX 1929, 1, A. Collart (IRSN); Bokoro, 20 III 1915, 1, R. Mayné (MRAC); Bolobo, 1941, 1, N'Kele (MRAC); Boma, 1, M. Tschoffen (MM); Bongide, 20 VIII 1957, 5 (FMNH); Botuna-Bokungu, 1950, 2, M. Boel (IRSN); Boyeka, Eala, 6 I 1915, 1, R. Mayné (MRAC); Buhunde-Matenda, 22 IX 1929, 1 (IRSN); Buhunde-Sawasawa, 15 IX 1929, 3, A. Collart (IRSN); Bumbuli, 1 IV 1915, 3, R. Mayné (MRAC); Bunia, 1939–1940, 3, F.F. Maristes (MRAC); Bunie, 29 X 1956, 1 (FMNH); Buta, 3, S.M. Reine Elisabeth (MRAC), 25 VI 1941, 1, Fr. Hutsebaut (MRAC); Congo da Lemba, V 1912, 1, XII 1912, 7 + 1 ab. baumanni , I 1913, 2, R. Mayné (MRAC); Coquihatville, 12 III 1911, 16, L. Burgeon (MRAC); Djombo, X 1912, 3, R. Mayné (MRAC); Dorsale de Lubero, Mt. Muteke, VI–VII 1963, 1 ab. baumanni, M.J. Celis (MRAC); Dungu, X 1919, 2 ab. baumanni , XI 1919, 1 + 1 ab. baumanni , III 1920, 1 ab. baumanni, P. Van den Plas (MRAC); Eala, IX 1912, 3, V 1914, 1, I–II 1917, 1, R. Mayné (MRAC), IV 1931, 7, H.J.Bredo (MRAC), I 1936, 42, J. Ghesquière (MRAC); Eala-Bokatola-Bikoro, IX– X 1930, 1, P. Staner (MRAC); Elisabethville, XI 1930, 1, R. Massart (MRAC); 1957–1958, 1, Ch. Seydel (MRAC); Equateur, 1925, 1, J. Berteaux (MRAC); Equateur, Bokuma, XII 1951, 5, I–II 1952, 1, III 1952, 2, VII 1952, 16, III 1954, 1, Lootens (MRAC); Equateur, Botende n. Yolombo, X 1922, 1, Hulstaert (MRAC); Faradje, Missa, 22 II 1930, 1, A. Collart (IRSN); Galli-Koko, Kasai, 6, R. Carlier (MRAC); Gandajika, 1947, 1 ab. dissoluta, P. Henrard (MRAC), 5 X 1949, 6 ab. dissoluta, 10 VIII 1950, 21 ab. dissoluta, 25 VIII 1950, 1 ab. dissoluta, 12 IX 1950, 13 ab. dissoluta, De Francquen (MRAC); Haut Tshuapa, Ikela, 1936, 7, Buckinckx (MRAC); Haut Tshuapa, Yolo, IV 1937, 1, Buckincx (MRAC); Haut Uele, Abarambes, 1 (IRSN); Haut Uele, Abimva, VI–VII 1925, 1 ab. baumanni, L. Burgeon (MRAC); Haut Uele, Moto, 1920, 2, 1922, 1, 1923, 6 + 1 ab. baumanni, L. Burgeon (MRAC); Haut Uele, Tuku, IV 1915, 1 ab. baumanni, Van der Plas (MRAC); Haut Uele, Watsa, XI 1913, 1, XI 1919, 1, 1922, 1 ab. baumanni, L. Burgeon (MRAC); Ikenge, 18 IX 1912, 7, 1 IV 1914, 3, R. Mayné (MRAC); Inongo, 1 III 1915, 1, R. Mayné (MRAC); Irangi, IX 1992, 2, X 1992, 3, H. Hinkel (TW); Irumu- Mombasa, 14 X 1932, 4, L. Lebrun (MRAC); Ilenge, I 1918, 2, R. Mayné (MRAC); Itoka, X 1912, 40, R. Mayné (MRAC); Ituri, Bunia, VI 1938, 1 + 8 ab. baumanni, Lefèvre (MRAC); Ituri, La Moto, Madyu, 1 ab. baumanni, L. Burgeon (MRAC); Ituri, Medje, IV 1914, 1, Chrusty (MRAC); Kabambare, 1, Lt. Delhaise (MRAC); Kabare, IV– VI 1952, 1 ab. baumanni, S. Vandevelde (MRAC); Kahuzi-Biega Park, VIII 1984, 3, L. Bartolozzi (MZUF); Kahuzi, Tshiranga, V 1993, 1 ab. baumanni, Ch. Neinhuis (TW); Kasai, Ipamu, 1922, 6, P. Vanderijst (MRAC); Kasai, Kondue, 1, E. Luja (MRAC); Kasai, Luebo, 1, G. Babault (MRAC), I–IV 1959, 1, P. François (MRAC); Kasai, Kwilu, II–III 1921, 1 ab. dissoluta, P. Vanderijst (MRAC); Kasai, Luisa, 1921, 1, L. Achten (MRAC); Kasai, Luluabourg, 30 I 1963, 1, J. Deheegher (MRAC); Kasongo, Niembo, 15 V 1918, 1, R. Mayné (MRAC); Katanga, Kabalo, III 1926, 1, H. Schouteden (MRAC); Katanga, Kikondja, V 1925, 1 ab. baumanni, de Witte (MRAC); Katanga, Kinda, 1, Don Cercle (MRAC); Kibali-Ituri, Kilomines, IX 1957, 1, C. Smoor (MRAC); Kibali-Ituri, Lodjo, VIII 1939, 113 (28 ab. baumanni ), Mme Lepersonne (MRAC); Kibali-Ituri, Nioka, I 1955, 1 + 1 ab. baumanni , XII 1955, 1 ab. baumanni, J. Hecq (MRAC); Kibombo, 1 XI 1910, 1, Bequaert (MRAC); Kikwit, 1920, 18, P. Vanderijst (MRAC); Kindu, 6 XI 1911, 2, L. Burgeon (MRAC), I 1924, 1, H. Lebeau (MRAC); Kisangani, XII 1970, 3, J. Taverniers (MRAC); Kisantu, 11, Vanderijst (MRAC, MM), 1931, 2, Vanderyst (MRAC); Kisenyi, 1 ab. baumanni, Loveridge (MCZC); Kivu, Beni Terr., M'Bau, 1200 m, 13 V 1953, 1, M. J. Celis (MRAC); Kivu, Butembo, IX–X 1965, 2 ab. baumanni, Célis (MRAC); Kivu, Butembo, vallée de la Musosa, V 1967, 1 ab. baumanni, Lejeune (MRAC); Kivu, Bwito, riv. Rwindi, VIII 1934, 1 ab. baumanni, Lt. Marlier (MRAC); Kivu, Goma, X 1932, 16 ab. baumanni, L. Burgeon (MRAC); Kivu, Irangi, 10–20 VI 1983, 1 ab. baumanni , 1–2 II 1986, 2, 18–21 I 1986, 1 ab. baumanni, A. Mühle (MD); Kivu, Kibati, n. Nyaragongo, 16 IV 1953, 1 ab. baumanni, J. Verbeke (IRSN); Kivu, Kisenje, IX 1907, 1 (ZMHU); Kivu lake, Kwidjwi Is., IX 1907, 2 ab. baumanni, Herzog Exp. (ZMHU); Kivu, Mabuita, XII 1935, 1 ab. baumanni, Boutakoff (MRAC); Kivu, Masisi, 1, A. Collart (IRSN); Kivu, Mulungu, 1939, 1 ab. baumanni, Hendricx (MRAC); Kivu, Rwankwi, VIII 1951, 1 ab. baumanni , IX 1951, 1 ab. baumanni, J. Leroy (MRAC); Kivu, Kabare Terr., Mushuere, 2 XI 1954, 1, N. Leleup (MRAC); Kivu, Katana, X 1932, 10 ab. baumanni, L. Burgeon (MRAC); Kivu, Kavumu ŕ Kabunga km 82, Mingazi, IV–VII 1951, 13, H. Bomans (MRAC); Kivu, Terr. Lubero, Mulo, 1960 m, VI–VII 1953, 1 ab. baumanni, J. Célis (MRAC); Kivu, Terr. Masisi, Mutakato, IX 1953, 3, N. Leleup (MRAC); N Kivu Lake, Rwankwi, IV 1946, 2, XI 1947, 1, XII 1947, 1, VIII 1951, 1, XI 1951, 1, VIII–XII 1951, 840 ab. baumanni, J.V. Leroy (MRAC); Kivu, Terr. Uvira, Itombwe, Hte Kambekulu, 2450 m, VI 1955, 2 ab. baumanni, N. Leleup (MRAC); Kivu, Tshibinda, XI 1932, 6 ab. baumanni, L. Burgeon (MRAC); Kivu, Vall. de la Ruzizi, Kanambo, III 1 959, 1 ab. baumanni, L. Benoit (MRAC); Kiwu-Ituri, 2 (LS); Kunungu, N'Kele, 1934, 1 ab. baumanni, H. Schouteden (MRAC); Kunzulu, IX 1917, 4, R. Mayné (MRAC); Kwilu, Kasai, II–III 1921, 1, P. Vanderijst (MRAC); Lac Leopold II, 11–24 VI 1925, 6, Prince Leopold (MRAC); Leopoldville, 8 VIII 1911, 1, A. Dubois (MRAC), 15 VII–8 VIII 1951, 1, P. Jobels (MRAC); Leopoldville, Sohal, 1, Ledrou (MRAC); Leverville, 1920, 1, P. Vanderijst (MRAC), 1928, 8 + 2 ab. dissoluta, Mme Tinant (MRAC); Libenge, 18 IX 1947, 5, 20 XI 1947, 2, R. Cremer and M. Neuman (IRSN); Lisala, 29 VII 1947, 1, Cremer & Neuman (IRSN); Lisala, Bokapo, 16 IX 1938, 13, J.J. Deheyn (MRAC); Lisala, Dua, 10 X 1938, 1, J.J. Deheyn (MRAC); Lisala, Dua-Ebola, 12–21 I 1938, 1 ab. baumanni, J.J. Deheyn (MRAC); Lisala, Ebola, 12–21 I 1938, 1, J.J. Deheyn (MRAC); Lisala, Modjuambe, 15 V 1938, 2, J.J. Deheyn (MRAC); Lisala, Momongo, X 1937, 4, J.J. Deheyn (MRAC); Lodima, VII–VIII 1906, 1, Waelbroeck (MRAC); Lokandu, 1939, 1, Lt. Marée (MRAC); Lokandu, Ile Blawa, VII 1939, 1, Lt. Vissers (MRAC); Lomami, Kamina, 1930, 1 + 23 ab. baumanni +1 ab. dissoluta, R. Massart (MRAC); Lomami-Kaniama, 1931, 1 ab. dissoluta, III–IV 1932, 15, R. Massart (MRAC); Lualaba, Kolwezi, III 1958, 2 ab. baumanni, V. Allard (MRAC); Lubero, 1937, 1 ab. baumanni, J. Ghesquière (MRAC); Lubutu, IX–X 1929, 4 (IRSN); Lubutu-Kirundu, 2 IX 1929, 1 (IRSN); Lubutu-Masua, 12 IX 1929, 2 (IRSN); Lubutu-Obongena, 7 IX 1929, 4, A. Collart (IRSN); Luebo, VIII 1921, 2, H. Schouteden (MRAC); Luhange, VI 1936, 1, A. Gonze (IRSN); Luluabourg, 1, P. Callewaert (MRAC), 14 IV 1939, 1 ab. dissoluta, 15 IV 1939, 3, 20 IV 1939, 1 ab. dissoluta, J. Deheyn (MRAC); Luluabourg, Katoka, 1938, 1, 1939, 1 ab. dissoluta, Vankerckhoven (MRAC); Lulua, Kabomba, XII 1937, 2, Carlier (MRAC); Lulua, Kapanga, XI 1932, 1 ab. dissoluta, II 1933, 1 ab. baumanni , III 1933, 3 ab. baumanni , I 1934, 1 ab. baumanni, G. Overlaet (MRAC); Lulua, Sandoa, X 1930, 1 + 1 ab. dissoluta, III 1931, 1, XI 1931, 1, F. Overlaet (MRAC); Lusambo, 1, M. Dupuis (MRAC); Lusambo et env., 1950, 3, P. Hostie (MRAC); Mahagi-Port, X 1939, 6 + 1 ab. baumanni, Mme Lepersonne (MRAC); Mandimbe-Uluku, 14 IX 1929, 1 (IRSN); Mandungu, 25 XI 1912, 44, R. Mayné (MRAC); Masisi, Uluku, 14 IX 1929, 2, A. Collart (IRSN); Masua-Lubutu, 9 IX 1929, 3 (IRSN); Mawambi, 1 (ZMHU); Mayidi, 1942, 1, 1943, 6 ab. dissoluta, Van Eyen (MRAC); Mayumbe, Bemba-Bungi, 3 III 1924, 1 ab. baumanni, A. Collart (MRAC); Mayumbe, Ganda-Sundi, 1915, 2, R. Mayne (MRAC); Mayumbe, Kiniati, 7–8 VI 1911, 2, R. Mayné (MRAC); Mayumbe, Seke Banza, 13 IV 1924, 8 + 6 ab. baumanni, A. Collart (MRAC); Mayumbe, Tshela, 13–27 II 1916, 4, R. Mayné (MRAC), 8 III 1 924, 1, A. Collart (MRAC); Mayumbe, Yingu, 7 XII 1923, 1 ab. baumanni, A. Collart (MRAC); Mayumbe, Zobe, 4–12 I 1916, 1, R. Mayné (MRAC); Mokoanghay, 1 ab. baumanni, L. Tilkens (IRSN); Mombali, VII 1913, 1, P. Vanderijst (MRAC); Mondombe, X 1912, 17, R. Mayné (MRAC); Mongbwalu, VII 1938, 25, Mme Scheitz (MRAC), VII 1939, 1, Mme Lepersonne, 22, Mme Scheitz (MRAC); Mongbwalu, Kilo, VIII 1937, 8, 1939, 38 +17 ab. baumanni, Mme Scheitz (MRAC); Mont Kabobo, terr. Albertville, Hte Kiymbi, 1700 m, IX 1958, 2 ab. baumanni , X 1958, 1 ab. baumanni, N. Leleup (MRAC); Moyen Kwilu, Leverville, 2, P. Vanderijst (MRAC); M'Paka, Terr. Libenge, VII–VIII 1959, 1, M. Pecheur (MRAC); Mpese, 5 VI 1937, 1, 26 VI 1937, 1, R.P. Cooreman (IRSN); Mulungu, I 1953, 1 ab. baumanni, R. Mayné (MRAC); Mutambo-Mukulu, 1936, 1 ab. baumanni, Richard (MRAC); Mwana n'Deke, V 1953, 2, De Francquen (MRAC); Nioka, 20 XI 1942, 1, J. Vrydagh (MRAC); Nyangwe, 26 XI 1910, 1, Bequaert (MRAC), 20 III 1918, 1, V 1918, 3, R. Mayné (MRAC); Poko-Nala-Rungu, 1912, 2, Me Hutereau (MRAC); Potopot n. Dima, 4 IX 1930, 2, de Witte (MRAC); Region de Sassa, 1895–1896, 5, Colmant (MRAC); Riv. Salonga, Bomputu, XII 1935, 2, J. Ghesquière (MRAC); Rutshuru, 26 IV 1936, 2 ab. baumanni, L. Lippens (MRAC); Rutshuru, Kilinga, 20 II 1936, 1 ab. baumanni (MRAC); Ruwenzori, N Alb.-Edw. See, II 1908, 1 ab. baumanni (ZMHU); Ruwenzori, NW Beni, I 1908, 1 (ZMHU); Ruwenzori, Fort Beni, I 1908, 3 (ZMHU); Ruwenzori, Mutawanga, 100–1300 m, X 1936, 1 ab. baumanni , XI 1936 – II 1937, 4, II–III 1 937, 4 (2 ab. baumanni ), Hackars (MRAC); Sanga, 26 VI 1926, 1 ab. baumanni, A. Collart (IRSN); Sankuru, Komi, I 1930, 1, 20 II 1930, 1, 22 II 1930, 1, 2 V 1930, 2, VI 1930, 1, J. Ghesquière (MRAC); Sankuru, Kondue, 1, E. Luja (MRAC); Sankuru, Lomela, V 1925, 6, J. Ghesquière (MRAC); Seke-Banza, Mayumbe, 13 IV 1924, 2, A. Collart (MRAC); Simba, Lokolenge, Lulonga, V 1927, 1, J. Ghesquière (MRAC); Stanleyville, 1924, 3, X 1925, 20, J. Ghesquière (MRAC); 19–21 IV 1928, 1, 27 VIII 1928, 1, 28 III 1928, 1, 13 IX 1928, 1, 31 V 1929, 3, VI 1929, 4, 13 VII 1929, 1, 23 XI 1929, 1, 1 XII 1929, 1, 21–31 XII 1929, 1, A. Collart (IRSN), X–XI 1949, 2, P. Waffelaert (MRAC); Stanleyville, Kilo, 3, L. Burgeon (MRAC); Tangayika, Moero, Nyunzu, I–II 1934, 2 ab. dissoluta, De Saeger (MRAC); Terr. de Dibaya, Kamponde, 1945, 3 ab. dissoluta, F. Allaer (MRAC); Thysville, 1 XII 1952, 3 ab. dissoluta, P. Basilevsky (MRAC); Tolo, XII 1913, 8, J. Maes (MRAC); Tshuapa, Bamanya, XII 1961, 1, V 1963, 1, 1–14 IX 1963, 1, II 1964, 1, IV 1964, 1, VI 1964, 1, IX 1964, 1, I–VI 1965, 1, Hulstaert (MRAC); Tshuapa, Boende, 1940/46, 2, Wynants (MRAC); Tshuapa, Bokuma, XII 1951, 2, III 1952, 2, IV 1952, 1, V 1952, 1, VI 1952, 1, VII 1952, 3, 1953, 2, 1954, 2, II 1954, 1, II–III 1954, 1, III 1954, 3, IV 1954, 1, XI 1956, 1, Lootens (MRAC); Tshuapa, Bokungu, 1949, 30, Dupuis (MRAC); Tshuapa, Etata, VII–VIII 1 969, 27, IX–X 1969, 27, V 1970, 1, J. Hauwaerts (MRAC); Tshuapa, Flandria, VIII–IX 1947, 1, Hulstaert (MRAC); Tshuapa, Ikela, 1955, 12, 1956, 29, X 1956, 10, XI 1956, 18, Lootens (MRAC, DBET); Ubangi, Gemena, 16 IX 1937, 1, L. Léontovitch (MRAC); Uele, Bambesa, 1937–38, 1, J. Vrydagh (MRAC); Uele- Itimbiri, Dingila, 26 IX 1932, 5, J. Vrydagh (MRAC), VI 1933, 1, H.J. Brèdo (MRAC); Uele, Kumu, 6 III 1938, 1, J.Vrydagh (MRAC); Uele-Pawa, 1938, 1, A. Dubois (MRAC); Upemba Nat. Park, Gorges de la Pelenge, 1150 m, 10–14 VI 1947, 1 ab. baumanni, de Witte (MRAC); Upemba Nat. Park, Kaziba aff. Senze aff. Lufira, 15–26 II 1948, 9 ab. dissoluta, de Witte (MRAC); Upemba Nat. Park, Kenbwale, Kalule N., 1050 m, 3–5 III 1949, 1 ab. dissoluta, de Witte (MRAC); Upemba Nat. Park, Kiamokoto-Kiwakishi, 1070 m, 4–16 X 1948, 3 ab. baumanni, de Witte (MRAC); Upemba Nat. Park, Lusinga, 1760 m, 18 XII 1947, 2 ab. baumanni, de Witte (MRAC); Upemba Nat. Park, Mabwe, Lac Upemba, 1– 12 VIII 1 947, 1 ab. baumanni , 18–27 VIII 1947, 1 ab. baumanni, de Witte (MRAC); Upemba Nat. Park, Mbunge-Bala, 1750 m, 25– 31 III 1 948, 2, de Witte (MRAC); Upemba Nat. Park, Mukana, 1810 m, 14 IV 1947, 3 ab. baumanni, de Witte (MRAC); Wamba, 1936, 4, Degrotte (MRAC); Wamba, Bayenga, 1 (MRAC); Wenga Ifomi, 1, E. Quineaux (IRSN); Yala, 3 XI 1912, 1, R. Mayné (MRAC); Yambata, 10 XII 1912, 5, R. Mayné (MRAC); Yangambi (Stan.), II 1939, 7, M. Brouwers (MRAC), XII 1939, 3, P. Henrard (MRAC), 10–19 VI 1948, 2, G. Benoit (MRAC), 18–25 VII 1952, 1, R. Mayné (MRAC), XI 1957, 1, P. Dessart (MRAC), XI 1959, 2, J. Decelle (MRAC); Yungu, 7 XII 1923, 1, A. Collart (IRSN).

EQUATORIAL GUINEA: Ayamiken, VIII 1958, 2, XII 1958, 4, II–III 1959, 13, III 1959, 61, XII 1959, 1, Sabater ( MCNB), I 1959, 11, B. Simon ( MCNB) ; Ayamiken-Bata, XI 1958, 4, Sabater (MCNB); Ayamiken-Rio Campo, XI 1958, 3, III 1959, 18, Sabater (MCNB); Ayamiken-Rio Benito, XI 1958, 1, II 1959, 5, Sabater(MCNB); Benito, 1 (CMNH); Benito env., Alén, 1– 15 VIII 1 906, 4, G. Tessmann (ZMHU); Mongo, 1946-1948, 86, J. Palau (MRAC), 1 IV 1946, 1, J. Palau (MCNB); Makome-Campagbt., 1–15 IV 1906, 1, G. Tessmann (ZMHU); Nkolentangan, XI 1907 - V 1908, 20, G. Tessmann (ZMHU); Sendye, II 1959, 4, B. Simon (MCNB); Sendye- Ayamiken, XII 1958, 13, XII 1958 - I 1959, 18, I 1959, 15, Sabater (MCNB); Sendye-Bata, II 1959, 1, Sabater (MCNB).

ETHIOPIA: Adis Ababa, Akakai river , 6 X 1980, 1, ab. baumanni, A. Demeter ( HNHM) ; Agheresalam , 7 VI 1963, 1 ab. baumanni, R. Linnavuori ( ZMUH) ; Bedel , III 1992, 1 (MS) ; Belleta Forest , 13–14 VI 1963, 9, R. Linnavuori ( ZMUH) ; Kaffa, Anderaccia, verso Ammaia, 1650 m, 26 X 1973, 1, Brignoli (DS), ponta naturale di Gurgutto , 26 X 1973, 1, Cottarelli (DS) ; Gemu-Gofa Prov., Arba-Minch , 1250 m, 29–30 V 1974, 1 ab. baumanni, G. de Rougemont ( MRAC) ; Ilubabor Prov., 15 km NW Ihora , 1600 m, VI 1973, 4 ab. baumanni, G. de Rougemont ( MRAC) ; Ilubabor Pr., Tépi, Masha , 8–9 V 2003, 1, B. Malec (FK); Jima, V 1992, 2 + 1 ab. baumanni, Trailin (RB); Kaffa, Bebeka, 10 V 1999, 1, K. Werner (MS); Kaffa, foresta a sud di Cossa, 2050 m, 24 X 1973, 1, Cottarelli (DS); Kaffa, Tra Gimma e Cossa, Torr. Aueddu del Diddessa e pozza, 1650 m, 24 X 1973, 3, Brignoli (DS).

GABON: Gabun , 2 ( ZMHU) ; Haut Ivingo, affl. Ogooué, 1906, 8, J. Gravot (MNHN); Lambarene, 1, A.C. Good ( CMNH) ; Libreville, 4 (SD); Libreville et env., V 1915, 1, G. Babault ( MNHN) ; Bas Ogooué, 1 (IRSN); Bas Ogooué, entre Lambarene et la Mer, 1901, 1, E. Hang (MNHN); Ogooué-Ivindo, 21 km road Mekambo to Madjingo, NE Zoula, 500 m, 26 XII 2000, 5, J.J. Wieringa (JW); Ogooué-Ivindo, Sassamongo, 630 m, 9 I 2001, 1, J.J. Wieringa (JW); Sud Ogowé , N'Gomo, 2, Junod ( MRAC) ; Ogove R., 5, Good (CMNH); Ogove R., Kangve, 16, A.C. Good ( CMNH) .

GHANA: Aschanti Region, Mampong , 28 IX 1971, 1, S. Endrödy-Younga ( HNHM) ; Asente Akem, Ashantis, 1, Junod (MRAC); Axim, 1, Besnard (MRAC); Kpandu , 13 III 1973, 1, W.F. Rodenburg ( NNML) ; Takoradi , 22 I 1967, 1 ab. baumanni , 18 VIII 1967, 1, 20 X 1967, 1, C. Besnard (BB); Volta Region, Amedzofe, 31 VIII 1967, 3, S. Endrödy-Younga ( HNHM) ; Western Region, Busua , 27 VIII 1969, 2, S. Endrödy-Younga ( HNHM) .

IVORY COAST: Forêt de Banco, 16 VIII 1966, 1, D. Thys and W. Verheyen ( MRAC) ; Bingerville , I 1964, 3, J. Decelle ( MRAC) ; Danane , I 1939, 1 ( MNHN) ; Mt. Tonkoui , 15–22 X 1973, 2, R. Linnavuori ( ZMUH) ; Orstom, 1964, 1, E. Binder (MHNG); Ton Koui Mt. n. Man , XI 1970, 2, E. Binder ( MHNG) .

KENYA: Kakamega Forest, 5100 ft., 18–22 I 1972, 1 (BMNH), 1 III 1976, 1, H. Gønget (MZK); Kitale, 2000 m, 24 I 1938, 1 ab. baumanni, A. Holm (NRS); Lembus Forest, Kapcholoi, 18 VIII 2004, 4 ab. baumanni , ABD (TD); Lembus Forest, R. Perakera Bridge, 15 VIII 2004, 1, ABD (TD); Lodwar, 20 XII 1995, 1 (DS); Masai mara, 13 V 1980, 1 ab. baumanni, D. Furth (PMNH); M’laba For., 2 II 1979, 4 + 7 ab. baumanni, T. Palm (LU); Monianku, Kisii, 24 VI 1978, 1 ab. baumanni, J.W. Waskevich (PMNH); Mt. Elgon, 1980 m, I–II 1979, 4, T.-E. Leiler (NRS); Nairobi, ICEPE, 1650 m, I 1999, 2 ab. baumanni, T. Wagner (TW); Victoria-Nyansa, Kwaa Kiloto to Kwa Muiya, III–IV 1894, 1 ab. baumanni, O. Neumann (ZMHU).

LIBERIA: Nimba, 12 IV 1964, 6 ab. baumanni, E. Binder ( MHNG); Mt . Nimba, 1400 m, 15 VIII 1966, 5 ab. baumanni, Ross & Lorentzen ( CAS) .

NIGER: Niger, 1 ab. baumanni, Benn. (ZMHU).

NIGERIA: Apeji, 2 ( BMNH) ; Lagos, VIII–IX 1948, 10, W. E. S. Merett ( BMNH, LS) ; Niger State, Mariga River, 80 km NW Minna , 11 XII 1987, 1, F. Kaplan ( TAU) ; Obudu Cattle Ranch , 27 VI 1973, 1 ab. baumanni, R. Linnavuori ( ZMUH) ; Old Calabar , 13 I 1900, 3, Annet ( BMNH) .

PRINCIPE IS.: Sao Thomé, 1 (MM).

REPUBLIC OF CENTRAL AFRICA: Fort Crampel, 15 + 4 ab. baumanni (IRSN); Fort Sibut, 1 (IRSN); Fort- Sibut, Oubangi-Chari, 1 (LS); Nola, 1 ab. baumanni (IRSN); Oubanghi-Chari, 2 + 1 ab. baumanni (MRAC).

REPUBLIC OF CONGO: Brazzaville, 1904, 1, J. Decorse ( MNHN) ; Loango, 2 (SD); Mongoumba, 1 (IRSN); Odzala Nat. Park , 400–500 m, 29 I–3 III 1 997, 2, S. Murzin & V. Siniaev (RB); Ouesso, 4, 2 III 1983, C. Besnard (BB); Samila chutes, 1, Mocquerys ( MRAC) ; San Benito R., 1885, 1, Gural (MNHN).

RWANDA: Bugesera, Biharagu, 27 II 1960, 1 ab. baumanni, N. Leleup (MRAC); Buschfang, 1800 m, 27 VII 1911, 1, H. Meyer (ZMHU); Cyangugu, Nyakabuye, 1–15 V 1983, 6 ab. baumanni , 24 IV 1985, 1 ab. baumanni , 12 V 1985, 3 ab. baumanni, H. Mühle (MD); Forêt Nyungwé, 9 XI 1985, 1 ab. baumanni, R. Jocqué (MRAC); Gihinga (Ruhengeri), 1800–2000 m, IX 1951, 1 ab. signaticollis, A.E. Bertrand (MRAC); Gishwati, Gakeri, X 1991, 1 (TW); Lac Mugesera, VIII 1954, 2 ab. baumanni, N. Leleup (MRAC); Nyakabuye Forest, 16 V 1987, 2 + 1 ab. baumanni (TW); Rubona, 4 IV 1963, 1 ab. baumanni , 5 IV 1963, 2 ab. baumanni , 7 IV 1963, 2 ab. baumanni , 8 IV 1963, 1 ab. baumanni, G. Pierrard (MRAC); Nyungwe, Kamiranzovu, 2000 m, X 1993, 3 ab. baumanni, T. Wagner (TW, DBET); Nyungwe, Pindura, X 1993, 1 ab. baumanni, T. Wagner (TW).

PRINICIPE ISLANDS: Sao Tomé, Mesquita , 29 III 1990, 1, Bordat (HPA).

SIERRA LEONE: Sierra Leone , 2 ( ZMHU).

SUDAN: Equatoria , Nimule, 11–13 III 1963, 2 ab. baumanni, R. Linnavuori ( ZMUH) ; Imatong Mts. , kateri- Gilo, 18 III 1963, 6 ab. baumanni, R. Linnavuori ( ZMUH) ; Imatong Mts. , near Gilo, 18–24 III 1963, 2 ab. baumanni, R. Linnavuori ( ZMUH) .

TANZANIA: Kirumba n. Muansa, 8 IV 1915, 1 ab. baumanni, Holtz ( ZMHU) ; Senga , 14 III 1915, 1 ab. baumanni, Holtz ( ZMHU) .

TOGO: Atakpamé-Bena, 12 IX 1973, 1, R. Linnavuori ( ZMUH) ; Bismarckburg, 20 IX–31 X 1890, 1, R.Büttner (ZMHU), 25 IX–12 XI 1892, 1 ab. baumanni, L. Conradt (ZMHU), 30 III–2 IV 1893, 1, L. Conradt (ZMHU); Misahöhe, Ende, III 1894, 1, E. Baumann (ZMHU); Missahoué, 650 m, VI 1963, 3, Y. Schach(MRAC); Oberssanga, Kag, 26 II–2 III 1913, 1, Tessmann (ZMHU); Uam Reg., Bosum, 21–31 III 1914, 1, Tessmann (ZMHU); Yob, V 1963, 1, Y. Schach (MRAC).

UGANDA: Ankole, XI 1919, 1 ab. baumanni, Loveridge (MCZC); Budongo Forest, 27– 28 VIII 1 971, 1 ab. baumanni , V 1972, 3 ab. baumanni , 14 XI 1972, 1 ab. baumanni, H. Gønget (MZK); Budongo Forest, Sonso, 6–16 X 2004, 7 ab. baumanni, T. Wagner (TW); Entebbe, X 1929, 1, G.A. Marshall (BMNH), I 1973, 1, H. Falke (MZSNV); Fort Portal, 15 km E Sebitoli, 1400 m, 23 XI–5 XII 1994, 8, M. Snižek (MS, DBET); Kalinzu Forest, Mpungu, 1200–1350 m, VI 1972, 1 ab. baumanni, H. Gønget (ZMC); Kampala, 15 IX 1915, 1, C.C. Gowdey (BMNH); Kampala, Tank Hill, 1300 m, 19 XII 1970, 1 ab. baumanni, H. Gønget (MZK); Kibale Forest, Toro Distr., XII 1972, 1 ab. baumanni , 19 II–3 III 1973, 5 ab. baumanni , 4 III 1973, 1, H. Gønget (MZK); Kisubi, 28 XI 1971, 1 ab. baumanni, H. Gønget (MZK); Kokanjero Mt., SW of Elgon, 6400 ft., 7–9 VIII 1911, 1, S.A. Neave (BMNH); Mabira, 1 ab. baumanni, R.A. Dummar (IRSN); Masindi Distr., Budongo Forest n. Sonso, 1°45’N, 31°35’E, 19–30 VI 1995, 9, 1–10 VII 1995, 3, 21–30 VII 1995, 1, 5–25 I 1997, 1, all ab. baumanni, T. Wagner (TW, DBET); Semliki Forest, 5–12 II 1997, 2 ab. baumanni, T. Wagner (TW); Mawaanga n. Kampala, 20 I 1973, 1 ab. baumanni, H. Gønget (MZK).

ZAMBIA: Abercorn, X 1943, 1 ab. baumanni, H.J. Brédo ( IRSN) ; Bangwelu Lake , Chiluwi Is., 26 XI 1946, 1 ( BMNH) ; Mniwilunga, II 1960, 1 (NMM). Mweru-Wantipa, I 1945, 1 ab. baumanni, H.J. Bredo (IRSN).

Figures

FIGURE 2. Distribution of Chiridopsis aubei.

FIGURES 20 – 23. Head. 20. Chiridopsis observabilis — species with flat clypeus; 21. Chiridopsis aubei — species with convex clypeus; 22. Chiridopsis tanganikana — species with long and angulate prosternal collar; 23. Chiridopsis tessellata – species with short and not angulate prosternal collar.

FIGURES 31 – 36. Chiridopsis aubei. 31 – 33. Nominotypical form, 31. Male dorsal, 32. Female dorsal, 33. Female lateral; 34 – 36. Variation of dorsal pattern.

FIGURES 37 – 42. Chiridopsis aubei. 37 – 38. Form described as ab. signaticollis, 39. Form described as ab. baumanni, 40 – 42. Form described as ab. dissoluta.

FIGURES 102 – 106. Chiridopsis quadriguttata. 102. Male dorsal, 103. Female dorsal, 104. Female lateral, 105 – 106. Variation of dorsal pattern.

FIGURES 43 – 49. Chiridopsis boutareli. 43. Male dorsal, 44. Female dorsal, 45. Female lateral; 46 – 47. Chiridopsis castaneoplagiata. 46. Dorsal, 47. Lateral, 48 – 49. Chiridopsis circe. 48. Dorsal, 49. Lateral.

FIGURES 24 – 30. Chiridopsis aequinoctialis. 24. Male dorsal, 25. Female dorsal, 26. Female lateral; 27 – 28. Chiridopsis convexula. 27. Dorsal, 28. Lateral, 29 – 30. Chiridopsis ruandana. 29. Dorsal, 30. Lateral.

FIGURES 17 – 19. Head. 17. Chiridopsis nigrostillata — species with broad clypeus; 18. Chiridopsis picturata — species with narrow clypeus; 19. Chiridopsis aequinoctialis — species with yellow clypeus.

FIGURES 80 – 86. Chiridopsis nigrosepta. Variation of dorsal pattern; 83 – 84. Chiridopsis nigrostillata. 83. Dorsal, 84. Lateral; 85 – 86. Chiridopsis klapperichi. 85. Dorsal, 86. Lateral.

FIGURES 94 – 101. Chiridopsis picturata. 94. Dorsal, 95. Lateral, 96 – 99. Chiridopsis tanganikana. 96. Male dorsal, 97. Male lateral, 98. Female dorsal, 99. Female lateral; 100 – 101. Chiridopsis tessellata. 100. Dorsal, 101. Lateral.

FIGURES 56 – 61. Chiridopsis observabilis. 56 – 58. Form described as Ch. observabilis, 56. Male dorsal, 57. Female dorsal, 58. Female lateral; 59 – 61. Form described as Ch. athinia, 59. Male dorsal, 60. Female dorsal, 61. Female lateral.

FIGURES 107 – 114. Chiridopsis parellina. 107. Dorsal, 108. Lateral, 109 – 110. Chiridopsis regionalis. 109. Dorsal, 110. Lateral; 111 – 112. Chiridopsis flavipennis. 111. Dorsal, 112. Lateral; 113 – 114. Chiridopsis zambiana. 113. Dorsal, 114. Lateral.