Pterostichus (Wraseiellus) crassiapex Shi & Sciaky

Shi, Hongliang, Sciaky, Riccardo, Liang, Hongbin & Zhou, Hongzhang, 2013, A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions, Zootaxa 3664 (2), pp. 101-135 : 111-112

publication ID	https://doi.org/ 10.11646/zootaxa.3664.2.1
publication LSID	lsid:zoobank.org:pub:105D178E-6DBF-401D-85BB-CDB041B67E91
DOI	https://doi.org/10.5281/zenodo.6154507
persistent identifier	https://treatment.plazi.org/id/03AA1008-5651-FFE2-C1D8-FD47FC60FCC1
treatment provided by	Plazi
scientific name	Pterostichus (Wraseiellus) crassiapex Shi & Sciaky
status	sp. nov.

Treatment

Pterostichus (Wraseiellus) crassiapex Shi & Sciaky View in CoL , new species

( Figures 7, 8 View FIGURES 7 – 12 , 23 View FIGURES 19 – 28 , 31 View FIGURES 29 – 34 , 59 View FIGURES 57 – 66 , 71 View FIGURES 67 – 84 , 96, 100 View FIGURES 98 – 106 , 107)

Type locality. Hubei: Badong County, Tiechanghuhang [=Tiechanghuang] (N30.75º, E110.30º), altitude about 1300 m.

Type materials. Holotype (IZAS): male, body length = 16.2 mm, board mounted, genitalia dissected and glued on plastic film pinned under specimen, " China, W Hubei, 27.VI–11.VII / road Badong-Yesanguan, 2003 / TIECHANGHUHANG / pit fall traps, 30.75 N 110.3 E / Jaroslav Turna leg. ~ 1300m "; " HOLOTYPE 3/ Pterostichus (Wraseiellus) / crassiapex new species / Des. SHI H.L. 2012" [red label] [ Figs. 7 View FIGURES 7 – 12 , 23 View FIGURES 19 – 28 , 31 View FIGURES 29 – 34 , 59 View FIGURES 57 – 66 , 71 View FIGURES 67 – 84 , 96, 100 View FIGURES 98 – 106 , 107]. Paratypes: 2 males (CRS), the same collecting data as the holotype but labeled as paratype.

Diagnosis. This new species can be distinguished from all the other species of this subgenus by the combination of the following characters: (1) 5th tarsomere with setae beneath; (2) pronotum cordiform, basal margin narrower than or subequal to the anterior margin ( Fig. 7 View FIGURES 7 – 12 ); (3) pronotum lateral margins with one seta on each side; (4) median lobe of aedeagus with apical lamella long, strongly bent to the left in dorsal view ( Fig. 31 View FIGURES 29 – 34 ); (5) right paramere long and bent, apex sharp ( Fig. 71 View FIGURES 67 – 84 B).

This species is most similar with Pt. andrewesi Jedlička from Jinfoshan. Besides their significant difference on male genitalia ( Figs. 29, 31 View FIGURES 29 – 34 , 67, 71 View FIGURES 67 – 84 ), these two species can be also distinguished by: (1) in Pt. crassiapex , lateral margins abruptly sinuate before hind angles, hind angles distinctly projecting ( Fig. 100 View FIGURES 98 – 106 ); in Pt. andrewesi , lateral margins gradually sinuate before hind angles, distance between the sinuation and hind angle superior than in Pt. crassiapex , hind angles slightly projecting ( Fig. 98 View FIGURES 98 – 106 ); (2) in Pt. crassiapex , elytral humeral teeth small but distinct; in Pt. andrewesi , elytral humeral teeth very small ( Figs. 98, 100 View FIGURES 98 – 106 ); (3) in Pt. crassiapex , punctures on proepisterna dense and coarse; in Pt. andrewesi , such punctures fine and sparse.

Description. Body length 15.0– 16.2 mm; dorsal side black, shining; tarsi and apex of tibiae brown; ventral side almost black. Males with elytral microsculpture transverse on the basal half, gradually turning to isodiametric near apex. Head. Frons with fine punctures restricted in the frontal furrows; antennae reaching elytra basal fifth; 3rd antennomere glabrous except apical setae; males with the terminal labial palpomere tubular. Pronotum cordiform, widest before middle, PW/PL = 1.29–1.32; lateral margins fully rounded near the middle, and abruptly sinuate before hind angles; hind angles distinct, rectangular or slightly acute, distinctly projecting; one mid-lateral seta present at about anterior third of lateral margins; basal fovea moderately deep, faintly defined; inner groove of basal fovea subparallel to median line, not curved to the outside, ended at a distance from the posterior margin of pronotum; outer groove of basal fovea somewhat longer than half length of the inner one, deep and reaching the basal margin of pronotum; ridge between outer groove and lateral margin distinct; area between inner and outer basal foveal grooves rugose and hardly convex; basal fovea and basal-median area strongly and densely punctate, sometimes basal-median area less so. Elytra with base slightly depressed between 3rd and 5th intervals; elytral shoulders moderately narrowed, humeral teeth small but distinct; intervals feebly convex; striae moderately deep, finely punctate or not; scutellar striae incomplete, forming short grooves, located between 1st and 2nd striae; umbilical pore series on 9th interval sparse in the middle, composed of 16–17 pores (6–7, 2, 7–8). Ventral side. Proepisterna and metaepisterna densely and coarsely punctate, proepisterna densely punctate; terminal sternum of males with a lunate ridge at about anterior third, slightly distinct and transverse, and slightly depressed after the ridge ( Fig. 96). Legs. 5th tarsomere with three or four pairs of setae beneath. Male genitalia. Median lobe of male genitalia bent less than 90 degrees, apex slightly bent to the ventral side and then the turned to the dorsal side ( Fig. 31 View FIGURES 29 – 34 A); ventral margin almost straight in the middle, dorsal margin gradually curved; apical orifice large, slightly turned to the left side ( Fig. 31 View FIGURES 29 – 34 B); apical lamella laminar, apex strongly thickened and turned to the dorsal side ( Figs. 31 View FIGURES 29 – 34 B, 59B); in dorsal view, apical lamella narrow and long, length about twice as the basal width, twisted to the left (the widest surface of apical lamella orientated to the dorsal-left side of median lobe), slightly turned to the left side ( Fig. 31 View FIGURES 29 – 34 B); apex of apical lamella round, slightly oblique to the left ( Fig. 59 View FIGURES 57 – 66 A). Right paramere fine and long, sickle-form, apex sharp ( Fig. 71 View FIGURES 67 – 84 B). Female unknown.

Distribution. Only known form the type locality, Tiechanghuang (Hubei) ( Fig. 110 View FIGURES 110 – 111 ).

Etymology. The name " crassiapex " is the combination of the Latin adjective etyma " crass -", which means "thick", and the Latin noun " apex ", referring to the median lobe of aedeagus with apex of apical lamella strongly thickened.

Remarks. Two females very close to this species were also studied by us: 2 females (IZAS), "Sichuan, Wushan, Liziping, 1850m " [ Fig. 8 View FIGURES 7 – 12 ]. But they are slightly different in the pronotum basal fovea (basal fovea only with very sparse punctures) and locality (about 60km NW of the type locality, but isolated by the Yangtze river gorge) from the holotype of Pt. crassiapex , so we didn’t included these two females in type series or determined them as this species, and only labeled them as " Pterostichus sp. near crassiapex ".