Campatonema lineata (Schaus) Sullivan, 2010

Sullivan, J. Bolling, 2010, New species of the Neotropical genus Campatonema Jones (Geometridae, Ennominae) with the first description of the female, ZooKeys 39 (39), pp. 263-272 : 269-270

Campatonema lineata (Schaus) comb. n.

Figs 4, 5, 8a, b, 10

Sabulodes ? lineata Schaus, 1911: 590 .

Type material. Syntypes: USNM. Type locality: Juan Vinas, Costa Rica.

Diagnosis. Campatonema lineata is readily separated from the other species of Campatonema by its wing pattern. In particular, no other species of Campatonema has broad, reddish-brown bands along the margins of both sets of wings.

Redescription. Male. Head – Palp short, less than length of round eye, slightly porrect, third segment 1/2 length of first two segments, which are equal in length. Palp mostly cream on inner surface and brown on external surface. Frons rectangular, brown. Area between antennae and scape cream. Antenna fasciculate with bristles originating in middle and distal end of each segment; antenna brown dorsally, two rows of scales per segment, distal row darker. Scales of collar erect, dirty cream in color. Thorax and abdomen – Dorsal thoracic scales cream with two tan spots on distal end. Abdomen cream with tan scales on posterior half of each segment, segments 6–7 with dorsal scales brown, tipped with cream, segment 8 tan with darker brown central spot. Ventral abdomen cream. Legs finely scaled with grey brown dorsally, cream ventrally, two pairs of tibial spurs on hind leg, one pair on middle leg. Wings – Forewing length 14 mm. Forewing slightly pointed, ground color cream heavily dusted with tan and chocolate brown; PM line originating subapically on costa and undulating to point at outer 1/4 of lower margin; PM line double on costal half, formed by two rows of chocolate scales with only inner one prominent and continuing to lower margin of wing. Traces of other tan lines parallel to wing margin both proximal and distal to PM line. Discal dot dark, distinct. Hindwing much as forewing, with a series of lines parallel to margin. PM line distad of discal dot darkest and most prominent. Margins of both wings tan with small chocolate patches at ends of veins. Ventral surface of wings with cream ground color and dusting of brown scales, becoming denser distally before forming a distinct marginal band. Apex of forewing lighter and forming a light band on forewing parallel to margin. Female. Slightly larger than male (forewing length 15 mm), maculation much like male but colors more intense. Antenna filiform. Male genitalia – Uncus rod shaped; gnathos arms indistinct, medial pad oval and covered with spines. Valves slightly shorter than in other species, more rounded apically. Area of bristles on valves adjacent to sclerotized costa and broadest apically. Annelar arms appear bifurcate, without sclerotized medial area. Arms of tegumen meeting medially below uncus, fused. Vincular arms straight. Furca deflected to left, extending to gnathos and with bristled area toward apex. Saccus squared. Juxta bending slightly to median at base. Aedeagus straight, pointed at apex, vesica expanded with patch of cornuti which areith cornuti slightly larger on basal side of patch. Pelt without distinct features. Female genitalia – Anal papillae elongated with apex rounded. Posterior apophyses twice as long as anterior ones, thin, straight, paddlelike at apex. Anterior apophyses with knob-like junction about 1/3 distance from posterior end, causing a slight redirection. Terminal ends paddlelike. Seventh tergite forming an oval shield with sclerotized ostial area below. Sclerotization on ostium forming a half collar dorsally. Ductus very short, forming a narrow neck to bursa, which expands slightly forming tear-drop shape. Signum on dorsal side a spiked sphere with hollow center. Pelt without distinct features.

Distribution and biology. Specimens have been collected in the provinces of Alajuela, Cartago, Guanacaste, and Puntarenas, Costa Rica, at elevations of 700–1500 m. Adults are on the wing throughout the year.