Hemicrepidius hemipodus ( Say, 1825 )

Etzler, Frank E., 2020, A Revision of the Genus Hemicrepidius Germar, 1839 (Coleoptera: Elateridae) of the New World, with Comments on Global Classification, The Coleopterists Bulletin (mo 18) 74, pp. 1-126 : 42-46

Hemicrepidius hemipodus ( Say, 1825)

( Figs. 5f , 6d , 19a–e , 20b )

Elater hemipodus Say 1825: 245 ; Say 1836: 176.

Asaphes hemipodus (Say) : LeConte 1853: 449; Candèze 1863: 210.

Diacanthus corporosus Germar 1843: 72 . [synonym by Candèze, 1863]

Hemicrepidius hemipodus (Say) : Horn 1880: 73 (as synonym of H. decoloratus ); Leng 1920: 170 (as synonym of H. decoloratus ); Becker 1979a: 410; Bousquet 1991: 183; Majka and Johnson 2008: 7; Majka et al. 2011: 139; Webster et al. 2012: 96; Bousquet et al. 2013: 184.

Athous aeneolus Melsheimer 1846: 156 ; LeConte 1853: 450 (as synonym of A. aereus ); Candèze 1863: 215 (as synonym of A. aereus ).

Athous aereus Melsheimer 1846: 156 .

Asaphes aereus (Melsheimer) : LeConte 1853: 450; Candèze 1863: 215; Horn 1880: 73 (as synonym of H. decoloratus ).

Diagnosis. Hemicrepidius hemipodus can be distinguished from all congeners by the following combination of features: An ear-like posterior end of the hypomeron so pronotal hind angle not visible in ventral view, and the curved prosternal process. Males can further be distinguished by having most punctures on the pronotal disc separated by one diameter or less, while both sexes can be distinguished by having lateral pronotal punctures subumbilicate and separated by <0.5× puncture diameter.

Hemicrepidius hemipodus could be confused with H. decoloratus and H. indistinctus . Key characters to separate H. hemipodus and H. decoloratus are provided in the diagnosis for H. decoloratus above. With H. indistinctus , only female H. hemipodus would likely be confused. They can be separated by female H. hemipodus having much closer and subumbilicate punctures on the lateral side of the pronotum, the punctures on the head being very dense and close, and in most by the slight metallic reflection on the elytra.

Redescription. Length 10.0–16.0 mm, width 2.0–4.0 mm; color uniformly brown to black or with elytra brown to dark brown and paler than dark brown to black pronotum, legs and antennae variable, yellow-brown to dark brown, antennae in most darker than legs, sometimes edges of pronotum, edges of elytra, and edges of abdominal ventrites paler; integument shining, some with slight metallic reflection. Setae moderately long, dense on pronotum in most, shorter on elytra and ventral surfaces, depressed, yellow or gray. Head: Punctures subumbilicate, nearly contiguous; frons weakly concave anteromedially when viewed dorsally; frontal carina broadly rounded when viewed from the front, often interrupted medially by punctation, frontoclypeal region complete and narrow medially, weakly projecting in lateral view; eyes moderate, rounded, somewhat bulging, ocular index 65–74 (n = 5); antenna weakly serrate, antennomere 2 subcylindrical, antennomere 3 subcylindrical, 1.5–2× length of 2 and with similar punctation to 2, antennomere 3 subequal in length to 4, antennomeres 4–11 about 3× longer than wide, becoming longer and narrower apically, antennomere 11 oval, sometimes with tip subtly constricted, antenna extending beyond pronotal posterior edge by 1.5–2.5 antennomeres. Prothorax: Pronotum as wide as long to 1.2× wider than long, widest at hind angles (most males) or at midlength (females); sides weakly to broadly arcuate, gradually narrowing anteriorly; hind angles divergent or weakly divergent, lateral edge sinuate anterior of hind angle, dorsal carina variable, absent (some males) to present, if present, thin or just slightly narrower than lateral carina (most females), parallel to lateral carina; punctures appearing simple, nearly touching to separated by one to two puncture diameters on disc, many females less punctate than males, punctures larger, shallower, and closer laterally, nearly contiguous, subumbilicate; posterior edge with sublateral notches. Hypomeron with punctures similar in size to lateral pronotal punctures, subumbilicate, nearly contiguous anteriorly and further apart posteriorly, small impunctate area present on posterior one-fifth at most; posterior margin ear-like ( Fig. 19c ) such that hind angle not visible in ventral view. Prosternum with punctation smaller than punctation on hypomeron, appearing simple, punctures nearly contiguous laterally, further apart medially, separated by two puncture diameters or more; anterior lobe slightly deflexed, slightly truncate anteriorly and partially covering ventral mouthparts; prosternal process slightly curved dorsad between procoxae in lateral view, narrow and straight past procoxae. Pterothorax: Mesoventral cavity with lateral edges sinuate, not visible anteriad mesocoxae in lateral view; posterior edge of cavity truncate. Metaventrite with punctures smaller than punctures on hypomeron, simple, dense laterally, nearly touching, becoming less dense medially. Scutellar shield elongate, about twice as long as wide, broadly rounded posteriorly, sides weakly sinuate, dorsal surface weakly convex, anterior margin nearly straight, setae radiating from midline, longer than setae on elytra. Elytral striae weakly impressed basally; interstriae flat to slightly convex, punctures moderate in size, scattered, giving wrinkled appearance; basal carina simple. Legs with lobes on tarsomeres 2 and 3 moderately wide, visible dorsally, lobe on 3 just slightly larger than lobe on tarsomere 2, lobe on tarsomere 1 small, lobe on tarsomere 4 smallest, both lobes on tarsomeres 1 and 4 visible in lateral view. Abdominal ventrites: Punctures simple, slightly smaller than punctures on metaventrite, nearly contiguous, less dense medially; fifth ventrite narrowly rounded; sides of ventrites 1 to 4 with microserrations partially obliterated in most, limited to anterior half of each ventrite with posterior half a series of shallow pits, or only noticeable at highest magnification (56× here), microserrations on ventrite 5 appearing absent. Male genitalia ( Figs. 19d, e ): Parameres broadly sinuate laterally and medially, pre-apical expansion acutely pointed laterally, apex of paramere broadly rounded, membranous, setose ventral patch near pre-apical expansion and 4–5 dorsal apical setae. Median lobe moderately thick, slightly sinuate laterally, basal struts broadly curved laterobasally. Basal piece with membranous area broadly curved medially in ventral view.

Distribution. Specimens were examined from the following states: UNITED STATES: AR, AZ, CT, DC, FL, IA, IL, IN, KS, LA, MA, MD, MI, MO, MS, NJ, NM, NY, OH, PA, SC, TN, VA, WV, WI ( Fig. 20b ). Canadian records for this species (e. g., Bousquet et al. 2013) likely refer to H. decoloratus and the status of this species in Canada needs confirmation.

Notes. This species has been historically confused with H. decoloratus . Areas of confusion with that species are discussed under notes for H. decoloratus above.

Athous aeneolus and A. aereus have been synonymized with H. hemipodus . These names have long been associated with the hemipodus-decoloratus complex, and are best placed here. It appears that Elater hemipodus was described from a female specimen, and females are generally more convex and less punctate on the disc than males (compare Figs. 19a and 19b ). Males match the type of A. aereus in the Melsheimer collection in the MCZC. The amount of sexual dimorphism observed in this species is common within Hemicrepidius .

The one remaining troubling name is A. aeneolus . LeConte (1853) stated that A. aereus is the female form and A. aeneolus is the male form of the same species. However, the specimen labeled A. aereus in the LeConte collection is actually a male, not a female. Melsheimer (1846), however, mentioned both sexes in his description of A. aereus , suggesting this species was described from more than one specimen and both sexes. Adding to this issue is that all six specimens in the Melsheimer collection under the name A. aereus are males, likely due to label removal or transfers over time. This means the original types of these two names are likely mixed within this series of six specimens. The type of A. aeneolus is considered lost, as it is impossible to determine which ones were examined by Melsheimer at the time of description.

As with most Say names, the type of E. hemipodus is either lost or a specimen identified by Say with his identification label, and a potential neotype, is located in a collection somewhere in Europe ( Prena 2018). The original description of this species closely matches two specimens labeled with the species name in the LeConte collection in the MCZC. Since the designation of a neotype of H. decoloratus here should eliminate confusion between these two species, I have not done so here until material in European collections can be examined thoroughly.

A series of three specimens labeled Diacanthus corporosus are located in the Museum für Naturkunde, Berlin, Germany (B. Jaeger, in litt.). One of the three has a lectotype label from M. C. Lane (unpublished, here desiginated) and is a typical female of this species .

Types Examined. Of Elater hemipodus : Type lost or missing; type locality Pennsylvania [?]. Of Diacanthus corporosus : Lectotype (here designated): 16536/ corporosus Kn. Grm., Am. Sept. Kn. / Lektotypus Nr.- / Lectotype Diacanthus corporosus M. C. Lane 1964 / Hist.-Coll. ( Coleoptera ) Nr. 16536 Corymbites corporosus Germ., Americ. sept. Knoch, Zool. Mus. Berlin; type locality North America, not further limited; in ZMHB. Of Athous aeneolus : Type not located, likely lost; type locality Pennsylvania; in MCZC[?]. Of Athous aereus : Lectotype (here designated): Red label, Melsheimer specimen #1; male; type locality Pennsylvania; in MCZC (lectotype designation unpublished by Becker).

LeConte Collection (5, MCZC). [pink disc]/ “ A. aereus (Mels) !/ decoloratus #6; [pink disc]/ var. aeneolus Mels. !/ decoloratus #7; [pink disc]/ decoloratus #9; [pink disc]/ Asaphes hemipodus (Say) Lec. , corporosus Germ. / decoloratus #10; [pink disc]/ hemipodus (Say) Mels / decoloratus #11.

Horn Collection (6, MCZC). In H. hemipodus tray: Pen., #1; [pink disc], #3; [yellow disc], #4. In H. decoloratus tray: N.H., #6; N.H., #7; no locality, #8.

Melsheimer Collection (11, MCZC). With A. hemipodus label, six specimens without locality labels. With A. aereus type, five specimens without locality labels.

Ziegler Collection (1, MCZC). Pa.

Specimens Examined (271). UNITED STATES: Coll. Janson, Ex. Candèze / Asaphes aereus Mels, Est. Un. LeC. [one with pink disc, purple rim] (2, BMNH); [red rectangle]/ [yellow rectangle] 494/ Athous aeneolus Mels., Am. Bor ./ Coll. Janson, ex Deyrolle (1, BMNH). Arizona: [Graham Co.], Graham Mts. , May-4-1964 , K. Stephan leg. (1, FSCA). Arkansas: Marion Co., 1897/ May (1, OSAC0000569393 ). Connecticut: [Hartford Co.], Southington, 7-5 (1, MCZC); New Haven Co. , Hamden , Broosvale Park , 1-VII-1973 , leg. K. W. Brown (1, PMNH); New Haven, West Rock nr. Judges’ Cave , 19-VI-1973 , K. W. Brown (1, PMNH). District of Columbia: N. Amer. / 4.74, E. Doubleday , decoloratus det. Horn (1, BMNH). Florida: Alachua Co., 5-7-1959 , W. R. Cave (1, FSCA). Illinois: N[orthern] Ill [inois], June [18]83 (5, WSUC); Champaign Co , 23-VI-1926 , K. F. Auden (1, INHS830408 ); Champaign , June-17-1939 , coll. C. O. Mohr (1, INHS830417 ); Decatur , June-12- 1935 , Frison and DeL. (1, INHS830192 ); Coles Co., Fox Ridge St. Pk. , VII-5-1966 , John D. Pinte (1, CASC); Shobanier, June-16-[19]48, Sand and Becker / yucca flowers (1, INHS830341 ); Urbana (1, INHS830294 ); Urbana , 6-7-1927 , A. E. Miller, coll. (1, FSCA); Urbana , 6-25-1940 , W. V. Balduf (1, INH830209 ): Urbana , June-20-1941 , J. Slater (1, INHS830253 ); Urbana , June-24-1948 , H. H. Ross (1, INHS830328 ); Vermilion Co., 27-VI-1926 , K. F. Auden (1, INHS830420 ); Wheaton , June-27-29-1952 , M. H. Hatch (1, OSAC0000541751 ); [Wabash Co.], Allendale , June-7-1941 , Ross and Mohr (1, INHS830525 ); Wills [sic] Co., Lockport , VI-28- 1961 , N. G. Seaborg (1, CASC); [247]/ Hart Coll’ n (1, INHS830423 ); [5908]/ Hart Det (1, INHS830333 ); [6124] (1, INHS830396 ); [6126] (1, INHS830442 ); [2(?)6] (1, INHS830419 ); [3442] (1, INHS830340 ); [18215] (1, INHS830338 ); [18257] (3, INHS830331 , INHS830337 , INHS830339 ); [18615] (1, INHS830330 ); [38826] (1, INHS830507 ). Indiana: Allen Co., New Haven , 7-17-1981 , N. M. Downie (1, FSCA); [Marion Co.], Camp Belzer, BSA, Indianapolis , July-15-1966 , Charles E. White (3, FSCA); Indianapolis, 7-14-[19]10 (1, MCZC). Iowa: [Des Moines Co.], Burlngtn (1, MCZC); [Story Co.], Ames , 6-9-[19]39, W. W. Wirth (1, LSAM). Kansas: Crawford Co. , 3mi NE Pittsburg, 37°26.64’N, 94°37.04’W, 1-13-VI-2005 , G. A. Salsbury, ex canopy trap, KAN1S05 098 (1, SEMC, SM0695929 KUNHM-ENT); Pittsburg , 2 mi W, 37°25.16’N, 94°45.47’W, 6-15-VI-2006 , G. A. Salsbury, ex canopy trap KAN1S06 064 (1, SEMC, SM0806259 KUNHM-ENT); Douglas Co., F. H. Snow (2, SEMC); June-17-1919 , William F. Hoffman (1, SEMC); Lawrence , 6-20-1926 , W. Benedict (1, SEMC); Jefferson Co., University of Kansas Field Station unit 3001, Rockefeller Prairie , 39.04529°N, 95.20491°W, 13-VI-2008 , Z. H. Falin, ex fogging crowns/corms of Tripsicum dactyloides KSR08 025 (1, SEMC, SM0794080 ); Johnson Co., 1915, R. H. Beamer (6, SEMC); Montgomery Co., Elk City S.P. Memorial Overlook area 37.27994°N, 95.77835°W, 11-VI-2010 , Z. H. Falin, K. T. Eldredge, ex. Sifted leaf litter, KDEE1F10 051 (1, SEMC0905888 ); Montgomery Co. State Fishing Lake , below dam, 37.16171°N, 95.68542°W, 9-11- VI-2010 , Z. H. Falin, K. T. Eldredge, S. R. Davies, J Ortega-Blanco, ex flight intercept trap, KDEE1F10 046 (2, SEMC0904689 , SEMC0904690 ); Neosho Co., IN Montana Bridge , 13-June-2010 , uv light, 37.28320, -95.10360, M. Bollin (3, SEMC1274794 , SEMC1274800 , SEMC1274814 ); Woodson Co., Woodson Co. State Lake W side, 25-V-1976 , leg. S. Hamilton, SWH9 US.KS.HamiltonS 1976.05.25 001 (1, SEMC1057562 ); Cross Timbers S. P., East Spillway access, nr lagoon, 37.73741°N, 95.92025°W, 8-11-VI-2010 , Z. H. Falin, K. T. Eldredge, S. R. Davies, J. Ortega-Blanco, ex flight intercept trap, KDEE 1F10 044 (1, SEMC0905227 ). Louisiana: [New Orleans Parish], New Orleans / Coll. Janson / decoloratus ID (1, BMNH). Maryland: [no data] (3, BYUC; 3, OSAC0000569386, OSAC0000569391 , OSAC0000569400 ); Allegany Co., Green Ridge St. For. , 22-VI-1985 , C. L. Staines, Jr. (1, FSCA); Baltimore , V-1902 / Van Dyke Collection (2, CASC); Baltimore , VI-6-[19]09, VI-12-[19]09 / F. E. Blaisdell collector (4, CASC); College Park, 5-2-[19]48, B. K. Dozier / Sweeping in woods (1, FSCA); College Park , 6-3-[19]32/ Collected by L. Putnam (2, WSUC); Little Orleans Twp , Little Orleans Campground, N 39.6305, W78.3890, sweeping, 6 June 2015, 4 June 2016, 3 June 2017 , Anthony Martin Deczynski (13, AMDC); Cecil Co., North East Twp, Rodney Scout Reservation , 25 June 2008 , Anthony Martin Deczynski (1, AMDC); [Charles Co.], Pope’ s Cr., May-22-[18]96 (2, MCZC); Montgomery Co., Bethesda. 11-VII-1984 , H. V. Weems, Jr., ultraviolet light (1, FSCA); Great Falls , 02-June-1987 , J. B. Stribling colr (1, MTEC, MTEC018134 [ Fig. 19c ]); Prince Georges Co., Laurel, Patuxent Wildlife Res. Ctr. , 28-June-1979 , W. E. Steiner (1, FSCA); Seneca, VI-19-[19]50, E. A. Fildes (1, FSCA). Massachusetts: [brown disc] [Cambridge 1871, Hubbard and Schwarz] (1, MCZC); Mt. Washington, June 22-25, 1913 (1, MCZC) ; Mt. Tom, July-4-[18]73 (1, MCZC). Michigan: Washtenaw Co, Ann Arbor , VII-18-1927 , N. K. Bigelow (1, FSCA). Mississippi: [Oktibbeha Co.], A & M college, 5-16-[19]31, C. F. Randle (1, WSUC); A & M College, 5-24-[19]31, J. H. Thomas (1, WSUC). Missouri: [no data] (1, BYUC; 1, MCZC); Henry Co. , 7mi E Clinton, 17- VI-1973 , G. H. Nelson / on Diospyros virginiana L. (2, FSCA); 14 mi SE Clinton , 28-VI-1973 , G. H. Nelson / on Diospyros virginiana L. (1, FSCA); Jackson Co., May-1951 , Ronald H. Pine (1, SEMC); Raytown , 16-VI-1977 , G. H. Nelson / ultraviolet light (1, FSCA); Newton Co., Shoal Creek at Alt. US Hwy 71, 24-V-1986 , C. B. Barr (1, LSAM); Ray Co., 5mi NW of Orrick on N, June-5- 1999 , coll F. W. Shockley, Black light trap (1, LSAM). New Mexico: [Colfax Co.], Kochler [Koehler?], VIII-9 (1, MCZC). New Jersey: [Camden Co.], S. Camden (1, MCZC); [Morris Co.], Schooleys Mt , VI-29-[19]11 / J. W. Green collector (1, CASC); Union Co., Cranford Twp, 7 Fifth Avenue , 28 June 2006, 4 June 2013 , Anthony Martin Deczynski (2, AMDC); as previous except, at light, 18 June 2014 (2, AMDC) ; Mountainside Twp, Watchung Reservation, 13-14 June 2007, Sylvio Codella and Anthony Martin Deczynski (1, AMDC). New York: [no data] (3, MCZC); Albany Co., Rensselaerville Huyck Preserve / Ross H. Arnett, Jr. collector, July-17-1975 (1, FSCA) ; [Greene Co.], Catskill, 6-19-[19]37 (2, CASC); Ontario Co., Geneva , V-30-VI-10-1991, VI-10-1991 , J. Huether (2, FSCA; DNA— MTEC023673 , MTEC023674 ); [Oswego Co.], Oswego, July-16- 1896 (1, MCZC) ; [Tompkins Co.], Ithaca, July-03, 7-July-[19]03, 12-July-[19]04, 17-July-[19]13, 18- July-[19]17, 20-July-[19]17, 15-July-[19]17, 27- July-[19]17, 28-July-[19]17, 25-July-[19]39 (4, CASC; 1, FSCA; 9, INHS 830446, INHS830448, INHS830456, INHS830458, INHS830462, INHS830468, INHS830469, INHS830475 , INHS830490 ); Ithaca, 1-July-[19]28, 18-July-[19]28/ H. A. Scullen (2, OSAC0000569392 , OSAC0000569398 ); Ithaca , VII-10-1940, VII-16-1940, VII-24-1940, VII-30- 1940 , J. N. Belkin (9, FSCA); Ithaca , 5-VII-1945, 9-VII-1947 , R. H. Arnett, Jr (4, FSCA); Ithaca , VII- 4-1966 , H. P. Lanchester (1, WSUC); Ithaca , 14- June-1995 , Michael S. Engel (2, SEMC [ Figs. 19a, b ]); Cornell Plantations , 18-June-1991 , M. F. Whiting (3, BYUC). Ohio: [no data] / Coll. Janson, ex Shaum / aeneolus ID (2, BMNH); Clark Co., Clark Lake, VI-16-[19]74, D. Brzoska (1, SEMC); Clinton Co. , VI-9-[19]55 / E. I. Hazard collector (1, FSCA); Delaware Co., Scioto River, south of O’ Shaughnessy Reservoir , 40.141°N, 83.120°W, 20-June-2016 , S. M. Clark (2, BYUC; DNA— MTEC023703 ). Pennsylvania: [no data] (3, MCZC); [no data] / Coll. Janson, Ex Candèze / hemipodus ID label (1, BMNH); No. 22 Reiche , N. Pennsylvania / Coll. Janson, Ex Dejean / A. decoloratus Say, Ath. viduus Dej., Amer. Bor. LeC (pars) (1, BMNH); [Cambria Co.], Ferndale , VII-25-1940 , Wm. Spector (1, FSCA); [Chester Co.], Avondale , VI-25- 1931 , coll. C. A. Thomas (1, CASC); [Cumberland Co.], Camphill, VI-2-[19]11/ Van Dyke collection (1, CASC); [Dauphin Co.], Harrisburg , VI-15-[19] 09/ Van Dyke Collection (1, CASC); Harrisburg , VI-20-[19]10 (1, MCZC); Hummelstown , VI-24- 1917, VIII-2-1919 / Van Dyke Collection (7, CASC); Del [aware]. Co., 6-2, 6-12 (2, MCZC); Villanova, VI-10-[19]70, D. Brzoska (2, SEMC); Franklin Co., Little Cove Road at Red Rock Road, N 39.72449, W78.05902, Sweeping , 3 June 2016 , Anthony Martin Deczynski (1, AMDC); [Monroe Co.], Water Gap (1, MCZC); [Montgomery Co.], Abington , VI-8 (1, MCZC); [Northampton Co.], Easton , VII-24-[19]11, VII-8-[19]17, VII-14-[19] 28 / J. W. Green collector (4, CASC); Easton, VI- 19-[19]21/ L. S. Slevin collection (1, CASC); Wind Gap, VI-25-[19]30, VII-2-[19]30, VI-11-[19]33, VII-7-[19]36, VI-11-[19]39, J. W. Green (6, CASC); [Philadelphia Co.], W. Park, 7-2/ Phila [delphia] (1, MCZC); Angora , VI-5 (1, MCZC); Manayunk (2, MCZC); Tioga Co. , Crooked Creek , 17-VI-1991 , J. Huether (2, FSCA, [ Figs.19d and 19e ]); [Westmoreland Co.], Jeannette, VI-30 (1, OSAC0000569395 ). South Carolina: [Richland Co.], Columbia , 13-V-1988 , E. M. Phillips (1, CESC). Tennessee: Blount Co., Cades Cove GSMNP, 13-VI-1986 , Howard V. Weems, Jr. (1, FSCA); GSMNP, Cades Cove ATBI Plot , Malaise Trap MT-0320010604, serviced 4- June-2001 , I. Stocks (1, LSAM0129231 ); GSMNP, Cades Cove ATBI Plot , MT-0320010618, 4-18- June-2001 , J. Burbank and R. Hightower (8, LSAM0128948 , LSAM0129076 , LSAM0129086, LSAM0129087, LSAM0129199, LSAM0129209, LSAM0129213 , LSAM0129215 ); GSMNP, Cades Cove ATBI Plot, Lindgren #03, 18-June-2-July-2001 , Parker, Stocks, and Peterson (1, LSAM0128802 ); GSMNP, Cades Cove ATBI Plot , 35°35’31”N, 83°50’17”W, Malaise Trap / 18-Jun-2-Jul-2001 , I. C. Stocks and R. Hightower, MT0320010702 (1, LSAM0103382 ); GSMNP, Cades Cove ATBI Plot , MT-042001010702, 18-June-2-July-2001 , R. Hightower and J. Burbank (1, LSAM0103565 ); GSMNP, Cades Cove ATBI Plot , Malaise Trap, MT- 0420020603, 9-May-3-Jun-2002 , J. H. Burbank (3, LSAM0134553 , LSAM0134558 , LSAM0134559 ). Virginia: Arlington Co., R. H. Arnett, Jr. (1, FSCA); Approx. 20 mi SW of Alexandria, 26-V-1975 / B. J. Harrington, P. S. Broomfield, B.M. 1975-392 (1, BMNH); [Fairfax Co.], Vienna, Je-13/ F. G. Carnochan (1, MCZC, MCZC-ENT 00642006 ); Vienna, J [un] e-13 (1, MCZC); [Fauquier Co.], Warrenton, 52 [?]-5-28, L. C. Woodruff (1, SEMC); Giles Co. , #164, Sinking Creek at Va Hwy 700, 17-June-1986 , 1950’, George W. Byers (1, SEMC); [Rockingham Co.], Blue Ridge Mounts and Shenandoah Valley , 3- 6-VI-1975 / B. J. Harrington, P. S. Broomfield , B.M .

1975-392 (1, BMNH). West Virginia: Grant Co., Cave Mountain, west of Landes , 8-VI-2001, S. M. Clark and R. A. Androw (13, BYUC; MTEC023704 ) ; [Marion Co.], Fairmont , 1928, P. N. Musgrove (3, CASC) . Wisconsin: [no data] (2, INHS830327 , INHS830344 ) . No locality info: (3, INHS830465 , INHS830479 , INHS830481 ; 1, MCZC) .

Figures

Fig. 5. Morphological characters of Hemicrepidius. Tarsi, lateral view: a) H. memnonius, b) H. bilobatus, scale bar between = 1 mm. Antenna types: c) Strongly serrate, with triangular antennomere 3 (H. niger), d) Serrate, with impunctate medial line (ml, arrow) of antennomeres (H. memnonius), e) Serrate, lacking medial line (H. californicus), f) Weakly serrate (H. hemipodus), all to scale, scale bar below d = 1 mm. Mesothorax: g) Lateral edges of mesothorax raised in near 90-degree angle, arrow (H. californicus), stepped line to highlight character, scale bar below body = 1 mm.

Fig. 6. Morphological characters of Hemicrepidius. H. memnonius: a) Mesothorax, ventral view, b) Crenulate basal carina on elytra (arrow), c) Detail of subumbilicate punctures, lateral edge of pronotum, e) Microserrations on sides of abdominal ventrites, arrow. H. hemipodus: d) Simple basal carina on elytra (arrow).

Fig. 19. Hemicrepidius hemipodus. Dorsal habitus: a) Male, b) Female. c) Pronotum, lateral view. Male genitalia:

Fig. 20. Distribution maps of examined Hemicrepidius specimens (blue/gray dots). a) H. falli, b) H. hemipodus.