Kaunia endyta (B.L.Rob.) King & Robinson (1980: 259)

Barreto, Jessica Viera & Sancho, Gisela, 2014, Typification of names in Kaunia (Asteraceae, Eupatorieae, Oxylobinae), Phytotaxa 188 (1), pp. 43-48 : 44-46

2. Kaunia endyta (B.L.Rob.) King & Robinson (1980: 259) . Basionym: Eupatorium endytum Robinson (1919: 13) .

Lectotype (designated here):— PERÚ. Puno: “between Sandia and the tambo Azalaya, on the way from Sandia to Chunchusmayo, 1500– 2000 m. alt.”, 5 June 1902, A. Weberbauer 1074 (Fragment in GH 00007650 [photo!]).

Robinson (1919) included in the protologue an explicit reference to the herbarium that houses the type of E. endytum : “Berlin, phot. and frag. Gr.”. The holotype was destroyed during World War II (1943), like the majority of the specimens held at B ( Sleumer 1949, Hiepko 1987). A photograph of this specimen at B (Field Museum, Photo negative 16249, http://emuweb.fieldmuseum.org/botany/search_berlin.php) was found. After consulting different herbaria (including B, MOL and USM), we could not find additional duplicates of the specimen collected by Weberbauer (number 1074). Here we designate the fragment at GH as the lectotype of E. endytum , according to Art. 9.12 of the International Code of Nomenclature for algae, fungi and plants ( ICN) ( McNeill et al. 2012), because this specimen was cited in the protologue.

3. Kaunia grossidentata (Hieron.) King & Robinson (1980: 259) . Basionym: Eupatorium grossidentatum Hieronymus (1908: 377) .

Lectotype (designated here):— BOLIVIA. Chuquisaca: “Camataqui, 2500 m.”, 10 February 1904, K. Fiebrig 3069 ( GH00007693 [photo!], isolectotypes S-R-8999 [photo!], LD 1061796 [photo!], G 00223766 [photo!], SI!).

See comments in Kaunia camataquiensis . The figure 1 A shows the photograph of the destroyed specimen at B (Field Museum, Photo Negative 16267, http://emuweb.fieldmuseum.org/botany/search_berlin.php) and the figure 1 C shows the chosen lectotype (http://plants.jstor.org/specimen/gh00007693?s=t).

4. Kaunia ignorata (Hieron.) King & Robinson (1980: 259) . Basionym: Eupatorium ignoratum Hieronymus (1908: 379) .

Lectotype (designated here):— BOLIVIA. “ Santa Cruz, 1600 m.”, May 1892, O. Kuntze s.n. ( NY 00169056 !).

Hieronymus (1908) described Eupatorium ignoratum on the basis of a specimen collected by O. Kuntze in Santa Cruz ( Bolivia). The holotype was destroyed at B, which housed the main collections of Hieronymus ( Stafleu & Cowan 1979). After consulting various herbaria, no further duplicates than that of NY were found, and then we proceed to designate the duplicate at NY as lectotype .

Robinson (1920) in “ The Eupatorium of Bolivia ” mentioned the specimen at NY and cited “alt. 2600 m.” perhaps by mistake .

5. Kaunia lasiophthalma (Grisebach) King & Robinson (1980: 259) . Basionym: Eupatorium lasiophthalmum Grisebach (1874:167) .

Holotype:— ARGENTINA. Tucumán: “ Cuesta de Siambón (inter Siambon et Juntas )”, March 1872, P. G. Lorentz 508 ( GOET 001519 [photo!], isotypes CORD 00006264 [photo!], CORD 00006265 [photo!]).

Recently , Freire & Ariza Espinar (2014) designated the specimen 508 collected by P. G . Lorentz and deposited in CORD ( CORD 00006264 ) as the lectotype of Eupatorium lasiophthalmum Grisebach. Probably , these authors were unaware about the existence of a duplicate of that collection at GOET . Dr. A . Grisebach was Professor at the University of Goettingen from 1837 and was appointed Director of the Botanical Garden there in 1875. A set of P. G. Lorentz collections from Argentina was kept at CORD and another set was sent to GOET where Dr. A . Grisebach had an opportunity to study the Argentinian material ( Hunziker 1960). It is unlikely that the specimen at CORD was studied by Dr. Grisebach. On the contrary, the specimen at GOET was undoubtedly studied by this botanist as can be confirmed by his handwritten note indicating the new name of the species in this collection (http://plants.jstor.org/specimen/ goet001519?s=t). Moreover , the locality indicated on the labels of the specimens of P. G . Lorentz 508 at CORD slightly differs from that indicated in the protologue (i. e. the labels at CORD point out “ Garabatal ” as part of the locality, instead of “ between Siambon et Juntas ” as indicated in the protologue and on the GOET specimen). Art. 9.19 of the ICN ( McNeill et al. 2012) requires that the author who first designates a lectotype must be followed, but that choice is superseded if (a) the holotype is rediscovered. Based on the information above and in agreement with Art. 9.19, we proceed to supersede the lectotype designated by Freire & Ariza Espinar since the holotype of E. lasiophthalmum is kept at GOET .

6. Kaunia longipetiolata (Sch. Bip. ex Rusby) King & Robinson (1980: 259) . Basionym: Eupatorium longipetiolatum Sch. Bip. ex Rusby (1893: 52) . Homotypic synonym: Ageratina longipetiolata (Sch. Bip ex Rusby) King & Robinson (1972: 92) .

Lectotype (designated here):— BOLIVIA: La Paz:“ Songo ”, November 1890, M. Bang 867 ( NY 00169096 !, isolectotypes BR 000000531285 [photo!], E 00433267 [photo!], GH 00273957 !, GH 00007783 [photo!], MO 714311!, NY 00169095 !, PH 00008376 [photo!], US 00610850 [photo!]).

Schultz Bipontinus (1865) first reported this taxon as “ Eupatorium longipetiolatum Sch. Bip. ” based on Mandon’s material (number 257), but this name was a nomen nudum. Rusby (1893) validly published the name indicating Bang’s specimen (“Songo, Nov. 1890 (867) = Mandon 257”) in the protologue, probably as a reference to the similarity between both specimens. In the NY herbarium, the institution to which H. H. Rusby was tightly related, there are two specimens collected by Miguel Bang with the number 867. One of them (number 867, NY 00169096) has the label of “PLANTAE BOLIVIANAE of Miguel Bang”, and a handwritten note indicating “Mandon nº 257 =” beside the number 867 of Bang’s collection. This handwritten note is similar to that in the protologue. The other duplicate collected by Bang (number 867, NY 00169095) does not have the handwritten note referring to the specimen Mandon 257. These two Bang´s specimens were treated as holotype and isotype respectively by John Pruski (annotation on the sheet, 1987) and as syntypes by Hind (2011). This last author assumed that the specimen Mandon 257 was cited as the other syntype by Rusby. For the sake of clarity, hereby we designate as lectotype the specimen Bang 867 ( NY 00169096) because it is well preserved and complete, and probably it is the very one that the author indicated in the protologue. Duplicates of Mandon 257 at BR, F, GH, MPU, NY, P, S, K and RB were further analyzed.

7. Kaunia saltensis (Hieron.) King & Robinson (1980: 260) . Basionym: Eupatorium saltense Hieronymus (1897: 786) .

Neotype (designated here):— ARGENTINA. “ Salta: Cafayate, Río Colorado ”, 8 January 1972, A. Krapovickas & C. L. Cristóbal 20687 ( F 1783063 !, isoneotypes CTES 00893 [photo!], GH 00273967 !, LP!).

In the protologue, Hieronymus (1897) cited two specimens with different localities, dates and collectors: “ Salta: bei der Stadt Salta ( LOR. u. HIERON., März 1873). Jujui: an nicht angegebenem Orte vermutlich bei der Stadt Jujui ( AUGUST STUMPF, Sommer 1874–1875)”. The herbarium B housed the main collections of Hieronymus ( Stafleu & Cowan 1979). We have found a photograph of the specimen collected by Lorentz and Hieronymus in Salta (Field Museum, Photo Negative 16355, http://emuweb.fieldmuseum.org/botany/search_berlin.php) and formerly kept at B. After consulting various herbaria including CORD and B, which hold important collections of specimens collected by Lorentz and Hieronymus, we could not find duplicates of the material cited by Hieronymus in the protologue. Due to apparently no original material is extant, we proceed to designate an appropriate neotype (Art. 9.7). The specimen collected by Krapovickas and Cristóbal (number 20687) at F is complete and well preserved, and agrees entirely with the characteristics indicated by Hieronymus in the protologue. In addition, the designated neotype has duplicates in other herbaria which assures the link of the type gathering to the name in case of the loss of the F material.