Syzygium nervosum

Soh, Wuu-Kuang & Parnell, John, 2015, A revision of Syzygium Gaertn. (Myrtaceae) in Indochina (Cambodia, Laos and Vietnam), Adansonia 37 (2), pp. 179-275 : 242-243

37. Syzygium nervosum A.Cunn. ex DC.

Prodromus Systematis Naturalis Regni Vegetabilis 3: 260 (1828). — Cleistocalyx nervosus (A.Cunn. ex DC.) Kosterm., Bulletin of the Botanical Survey of India 29: 17 (1987 publ. 1989), sphalm. nervosum , comb. illeg., non (Lour.) Blume (1850). — Cleistocalyx nervosum (A.Cunn. ex DC.) P.H.Hô, An Illustrated Flora of Vietnam (Câyco Viêtnam) 2: 63, fig. 3776 (1992). — Type: Roxburgh s.n. in Herb. Lambert 1816 (holo-, G-DC[microfiche E!]). (see Panigrahi & Mishra [1985] and Turner& Ghandi [2005] for detailed discussions on the typification; see Note 1 for discussion on S. angkolanum ). Eugenia operculata Roxb. [Hortus Bengalensis (1814) 37, nom. nud.]

Flora indica 2 [ed. 2, ed. W. Carey]: 486 (1832); Gagnep. in

Lecomte, Flore générale de l’Indochine 2: 817 (1920). — Syzygium

operculatum (Roxb.) Nied. in Engl. & Prantl., Natürlichen Pflan-

zenfamilien 3(7): 85 (1893), nom. illeg. — Cleistocalyx operculatus

(Roxb.) Merr. & L.M.Perry, Journal of the Arnold Arboretum 18:

337 (1937) [excl. Syzygium angkolanum Miq. ], nom. nov. pro

Syzygium nervosum DC. ; V.D.Nguyen, Medicinal plants of Viet-

nam, Cambodia and Laos: 282 (1993). — Type: Roxburgh s.n. in

Herb. Lambert, 1816 (lecto-, G-DC![microfiche], designated by

Panigrahi & Mishra [1985: 298]).

Eugenia paniala Roxb. [Hortus Bengalensis (1814) 37, nom. nud.] Flora indica 2 [ed. 2, ed. W. Carey]: 489 (1832). — Eugenia operculatus var. paniala (Roxb.) Duthie in Hooker, Flora of British India 2: 498 (1879), comb. illeg. — Cleistocalyx operculatus var. paniala (Roxb.) Chantaranothai & J.Parn., Kew Bulletin 48: 591 (1993). — Cleistocalyx nervosus var. paniala (Roxb.) J.Parn. & Chantaranothai, Novon 6: 201 (1996). — Cleistocalyx cerasoides var. paniala (Roxb.) I.M.Turner , The Gardens’ Bulletin Singapore 57: 26 (2005). — Type: Icones Roxburghianae no. 2255 (lecto-, K! here designated; isolecto-, CAL n.v.) (see Note 2).

Calyptranthes makal Blanco , Flora de Filipinas, ed. 1 (1837) 419, nom. illeg., non Raeusch. (1797). — Calyptranthes zuzygium Blanco , Flora de Filipinas, ed. 2: 293 (1845), nom. illeg., non Sw. (1788); Merr., Species Blancoanae: 288 (1918) [as synonym of Eugenia clausa C.B.Rob ]. — Type: Phillipines, Rizal Province, Antipolo, Sp. Blancoan. 0978 (neo-, US![00689480], here designated).

Eugenia cerasoides Roxb. , Flora indica 2 [ed. 2, ed. W. Carey]: 488 (1832). — Syzygium cerasoides (Roxb.) Raizada , The Indian Forester 84: 478 (1958). — Syzygium cerasoides Raizada , The Indian Forester 93: 755 (1967), nom. illeg. — Cleistocalyx cerasoides (Roxb.) I.M.Turner , The Gardens’ Bulletin Singapore 57: 26 (2005). — Type: Icones Roxburghianae no. 2256 (lecto-, K!, designated by Turner & Ghandi [2005: 25]).

Calyptranthes mangiferifolia Hance ex Walp., Annales Botanices Systematicae 2: 629 (1852). — Type: China, Macao, Hance Herb. (BM n.v.) (fide Merrill & Perry [1937: 337]).

Syzygium nodosum Miq. , Flora van Nederlandsch Indië 1(1): 447 (1855).— Type: Indonesia, Java, Palengang, Junghuhn s.n. (lecto-, L![U0004982], here designated; isolecto-, [L0012413]).

Eugenia operculata var. obovata Kurz, Forest flora of British Burma 1: 482 (1877). — [ Syzygium obovatum Wall., Wallich Numer. List 3552A & B (1831), nom. nud.]. — [ Syzygium wallichianum K.Presl, Abhandlungen der Königlichen Böhmischen Gesellschaft der Wissenschaften 5, 3: 500 (1845), nom. nov., nom. nud.]. — Syzygium nervosum var. obovatum (Kurz) A.Kumar, Journal of Economic and Taxonomic Botany 7: 664 (1985 publ. 1986). — Type: Bangladesh, Sillet, F.de Silva s.n., Wallich Numer. List 3552A (1831) (lecto-, K-W! here designated; isolecto-, E!, P!).

Eugenia holtziana F.Muell., Australasian Journal of Pharmacy 1: 199 (1886), sphalm. holtzei, in Systematic Census of Australian Plants 2: 101 (1889). — Type: Australia, Moritz Holtze 44 (holo-, MEL n.v.). (fide Hyland [1983: 108]).

Eugenia clausa C.B.Rob., Philippine Journal of Science, Section C Botany 4: 380 (1909). — Type: Philippines, Luzon, Province of Rizal, Bosoboso , Bur. Sci. 6760 (Robinson) (holo-, K n.v.; iso-, BR!).

Eugenia divaricatocymosa Hayata, Icones Plantarum Formosanarum 3: 118 (1913). — Type: China, Hainan, VI.1909, Konishi s.n. (holo-, TI n.v.) (fide Merrill & Perry [1937: 337]).

Eugenia suavis Ridl. , Journal of the Federated Malay States Museums 5: 160 (1915). — Type: Peninsular Malaysia, hill of Koh Samui, V.1913, Robinson 5730 (K!), syn. nov. (see Note).

[ Calyptranthes cuneata Buch. -Ham. ex Wall., Wallich Numer. List 3557B (1831), nom. nud. (E!, K-W!).]

[ Calyptranthes grandis Buch. -Ham. ex Wall., Wallich Numer. List 3554 (1831), nom. nud. (E!, K-W!)].

[ Calyptranthes tatna Buch. -Ham. ex Wall., Wallich Numer. List 3555 (1831), nom. nud. (E!, K-W!)].

[ Calyptranthes costata Buch. -Ham. ex Wall., Wallich Numer. List 3556 (1831), nom. nud. (E!, K-W!)].

[ Syzygium panealla Wall., Wallich Numer. List 3557A (1831) (K-W!, E!), nom. nud.].

[ Syzygium polyanthum auct. non (Wight) Walp. (1843): Thwaites, Enumeratio Plantarum Zeylaniae: 116 (1859), spec. C.P. 2801 (PDA n.v.)] (fide Kostermans [1981: 121]).

ADDITIONAL MATERIAL EXAMINED. — Vietnam. Ba Ria-Vung Tau: Ba Ria, IV.1861, Pierre s.n. (P). — Bac Kan: Ba Be National Park , 12.VII.2004, Daniel, N.T.Hiep & P.V.The DA 4752 ( HN) . — Cochinchina: Pierre s.n. ( BM, E, K). — Dong Nai: Bien Hoa , IX.1865, Pierre 56 (P). — Hanoi City: 30.III.1888, Balansa 2871 (P). — Kontum: Kon Plong, 16.XI.1988, Hien 366 ( HN) . — Lam Dong: 14.XII.1952, Schmid s.n. (P); 27.II.1953, Schmid s.n. (P). — Ninh Binh: 21.IV.1882, Bon 1515 (P). — Quang Ninh: Tien Yen, Hong Du forest reserve, VII.1918, 30.VI-3.VII.1918, Chevalier 37911 (P); Ha Coi, Tsai Wong Mo Shan and vicinity, 5.V-22.VI.1939, W.T.Tsang 29208 (P); Dam Ha, Sai Vong Mo Leng, 18.V-5.VII.1940, W.T.Tsang 29997 (P); idem, 8.V-5. VII.1940, W.T.Tsang 30053 (P); idem, 18.VII-9.IX.1940, W.T.Tsang 30343 (P). — Thanh Hoa: Cu Phach, 1885-1889, Balansa 2872 ( BM, K, L, P) .

DISTRIBUTION. — Widespread from Nepal to North Australia. In Indochina only documented in Vietnam and possibly not yet collected in Cambodia and Laos ( Fig. 29 ).

ECOLOGY OR HABITAT. — In forest and cultivated.

CONSERVATION STATUS. — IUCN Global Status: Least Concern (LC); IUCN Regional (Indochina) Status: Least Concern (LC).

PHENOLOGY. — Flowering: March, April, July, November, December. Fruiting: not documented.

VERNACULAR NAMES. — Vietnam: Diang (Dong Nai), Voi (Ninh Binh & Thanh Hoa), Voi ( Vietnam) .

USES. — The fruit is edible. The leaves are used as a beverage and are believed to be stomachic. A decoction of the leaves and flower buds is used as an antiseptic for washing boils and ulcers ( Nguyen 1993). The wood is used for house building ( Lemmens et al. 1995).

DESCRIPTION

Tree to 15 m tall. Bark greyish or brownish. Glabrous.

Figures

FIG. 29. — Distribution of Syzygium lineatum (DC.) Merr. & L.M.Perry (), Syzygium megacarpum (Craib) Rathakr.& N.C.Nair () and Syzygium nervosum A.Cunn. ex DC. () in Indochina.