Tetragonoderus quadriguttatus

Shpeley, Danny & Ball, George E., 2008, Taxonomic review of the Neotropical Tetragonoderus quadriguttatus assemblage (Coleoptera: Carabidae: Cyclosomini) with description of T. deuvei, new species, and new West Indian and Nearctic locality records, Insecta Mundi 2008 (50), pp. 1-16 : 7

Key to adults of the Tetragonoderus quadriguttatus assemblage and T. subfasciatus Putzeys

1. Elytron with discal setigerous punctures large, foveate; microsculpture mesh pattern markedly transverse (Fig. 3D and 4D); dorsal surface shining, subiridescent; preapical pale mark spotlike, in intervals 5-8 only, or fascia-like, in intervals 2-8 (Fig. 2D). Male genitalia with apical portion of phallus in dorsal aspect slightly spatulate (Fig. 5G), nearly flat. Geographical range: northern South America ( Venezuela) and Middle America ( Panamá) (Fig. 6) ........................... ......................................................................................................... T. subfasciatus Putzeys

- Elytron with discal setigerous punctures small; microsculpture mesh pattern isodiametric to distinctly transverse (Fig. 3A-C, and 4A-C); dorsal surface rather shiny or with sericeous luster; with two spots, preapical one in intervals 5-8 only (Fig. 2A, 2B and 2C), or unicolorous black. Phallus various (Fig. 4A-F). Specimen from South America (east of the Andes), West Indies or North America (Florida) ........................................................................................................... 2

2(1). Elytron with mesh pattern distinctly transverse (Fig. 3C and 4C), dorsal surface with more or less sericeous luster, and two-spotted. Phallus in dorsal aspect with apical portion short rather broad, apical margin broadly rounded (Fig. 5E), in lateral aspect projected dorsad (Fig. 5F). Geographical range: South America, from Paraguay northward to Colombia, and the West Indian islands of Grand Cayman, Jamaica, and Hispaniola ....... T. quadriguttatus Dejean

- Elytral microsculpture mesh pattern isodiametric (Fig. 3A and 4A) or slightly transverse (Fig. 3B and 4B); surface shiny, without sericeous luster, and two-spotted or uniformly dark. Phallus with apical portion flat (Fig. 5A-D), not projected dorsally. Geographical range: South America, principally south of Amazon Basin, and southern Florida ...................................................... 3

3(2). Elytron with mesh pattern mostly isodiametric, some sculpticells slightly transverse (Fig. 3A and 4A). Phallus with apical portion in dorsal aspect broadly triangular, tapered to obtusely pointed apex (Fig. 5A) ....................................................................... T. laevigatus Chaudoir

- Elytron with mesh pattern more transverse than above (Fig. 3B and 4B). Phallus with apical portion in dorsal aspect slender, curved slightly to left, apex narrowly rounded (Fig. 5C) ........ ............................................................................................................ T. deuvei , new species