Evagetes orientalis Lelej & Loktionov

Lelej, Arkady S. & Loktionov, Valery M., 2009, Review of the Evagetes crassicornis species-group (Hymenoptera: Pompilidae), with description of new species, Zootaxa 2230, pp. 42-50 : 45-47

Evagetes orientalis Lelej & Loktionov , sp. nov.

(Figs. 2, 6, 7, 10, 11, 14)

Diagnosis of male. New species is easily differentiated from other member of the E. crassicornis speciesgroup by short metaposnotum (ratio of metanotum length to metapostnotum length 3.3–4.5X in E. orientalis versus 1.7–2.9X in other species of this group), by the unique shape of S8. The volsella of the genitalia less swollen than in E. crassicornis , while it is not swollen in E. sahlbergi .

Diagnosis of female. New species is distinguished from other members of the E. crassicornis speciesgroup by short metaposnotum (ratio of metanotum length to metapostnotum length 4.0–5.4X in E. orientalis versus 1.8–3.7X in other species of this group).

Description of male. Body length 4.6–7.3 mm (holotype 6.2 mm). Fore wing length 4.0– 5.5 mm (holotype 5.5 mm). Head width 1.1–1.2X its height. Frons evenly convex. Ocelli small, POD: OOD 0.88– 1.13X. Ratio of genal median length to eye median length (lateral view) 0.25–0.45X. Malar space 0.22–0.45X pedicel width. Clypeus weakly longitudinally convex with weakly emarginated anterior border. Labrum flat with straight anterior border. Mandible tridentate. Flagellomere 1 length 1.22–1.38X its width. Relation of scape, pedicel and two first flagellomeres 24–30: 10–13: 16–20: 19–20. Mesosoma length dorsally 1.5X its width. Pronotum length 0.45X its width, posterior pronotal border angulate. Metanotum median length 3.3– 4.5X metapostnotum median length. Metapostnotum laterally shiny with transversal striae. Propodeum with sharply defined anterior horizontal and posterior vertical faces, its median length 0.8–0.9X its maximum width. Longer spur of mid and hind tibia not projecting over the apex of tarsomere 1. Fore claws bifid, asymmetrical, mid and hind claws symmetrical with additional inner tooth. Fore wing slightly infuscate, venation as in Fig. 10. Pterostigma small, second abscissa of R equal or longer than third abscissa of R. S8 (hypopygium) widened basally, distinctly emarginated apically, roof-like between basal carinae, with distinct short setae along lateral and apical border, and denser ones in apical and median parts (Figs. 6, 7). Genitalia as in Fig. 2.

Frons, temple, gena, supraclypeal area, propleuron with long scattered erect brownish setae. Mandible with two black strong setae and a few soft whitish setae. Fore coxa usually lacking setae, rarely with sparse short setae. Posterior half of pronotum, mesoscutum, mesoscutellum with sparse short setae. Mesopleuron, metanotum, propodeum, mid and hind coxa, and metasoma lacking setae. Lower part of face, clypeus, mandible basally with weak silver pubescence; frons, mesosoma, legs with pale-brownish pubescence; propodeum and T1 basally with silver pubescence, remainder metasoma with weak brownish pubescence. Body regularly micropunctate. Metasoma matt. Body and legs black, mandible black basally and brownish apically, T1 laterally and T2 baso-laterally ferruginous-red.

Description of female. Body length 6.5–9.1 mm. Fore wing length 5.4–7.1 mm. Head width 1.15–1.25X its height. Frons evenly convex. Ocelli small, POD: OOD 0.95–1.08X. Ratio of genal median length to eye median length (lateral view) 0.32–0.45X. Malar space 0.30–0.46X pedicel width. Clypeus longitudinally convex with weakly emarginated anterior border. Labrum flat with weakly emarginated or even straight anterior border. Mandible tridentate. Flagellomere 1 length 2.5–2.8X its width. Relation of scape, pedicel and two first flagellomeres 32–41: 13–15: 34–46: 31–42. Mesosoma length dorsally 1.3–1.4X its width. Pronotum length 0.4X its width, posterior pronotal border angulate. Metanotum median length 4.0–5.4X metapostnotum median length. Metapostnotum laterally shiny with transversal striae. Propodeum with sharply defined anterior horizontal and posterior vertical faces, its median length 0.8–0.9X maximum width. Tarsal comb short, apical spine of tarsomere 1 not touching the apex of tarsomere 2, apical spine of tarsomeres 2 and 3 projecting over the apex of tarsomeres 3 and 4 correspondently, tarsomere 1 with three spines, tarsomere 2 with two, tarsomere 3 with one, tarsomeres 4 and 5 without spines. Longer spur of mid tibia 0.75– 0.80X tarsomere 1, longer spur of hind tibia 0.67–0.74X tarsomere 1. Claws symmetrical with additional inner tooth. Fore wing slightly infuscate, venation as in Fig. 11. Pterostigma small, equal or slightly longer than first abscissa of R. Second abscissa of R 1.8–3.0X third abscissa of R.

Frons, supraclypeal area with sparse short erect black setae. Fore coxa with scattered long black erect setae, gena, propleuron with denser setae. Mandible with two long curved strong black setae and one-three additional shorter ones. Posterior half of pronotum, mesoscutum, mesoscutellum with sparse short brown setae. Mesepisternum anterior to coxa lacking setae or with a few short black setae. Metanotum and propodeum lacking setae. Posterior border of S3–S5 with sparse erect black setae, S6 with scattered setae. Head, mesosoma, coxae ventrally with brownish pubescence. Mid and hind coxa posteriorly with silver dense pubescence. Propodeum with sparse iridescent pubescence. Terga with sparse pale pubescence in basal half and brownish pubescence in apical half. Body regularly micropunctate. Body and legs black, outer eye orbit near vertex with small pale patch, rarely inner orbit with very small pale patch also, mandible brown basally, yellow-orange medially and brownish apically; T1 apically and T2 basally ferruginous-red.

Holotype data: ď, Russia: Primorskiy Terr., Anisimovka, 15.VI.1997 (A. Lelej) [ IBSS] ( Fig. 14 ).

Paratype data: Primorskiy Terr.: 1 ď, Partizansk, Chandalaz Mts., 26.IX.1974 (A. Berezantsev); 1 ď, Khasan, 26.VII.1977 (A. Lelej); 1 Ψ, Tikhookeanskiy, 26.VIII.1978 (A. Lelej); 4 Ψ, 20 km W Kamen- Rybolov, 5–8.IX.1978 (A. Lelej); 1 ď, Sergeyevka, 21.VII.1979 (S. Belokobylskij); 1 ď, 30 km E Spassk, 16.VI.1980 (S. Belokobylskij); 1 ď, 20 km NE Spassk, 24.VII.1998 (S. Belokobylskij); 2 Ψ, Anisimovka, 4.IX.1982 (A. Lelej); 2 ď, the same place, 3, 5.VIII.1983 (A. Lelej & E. Budris); 1 Ψ, Novogeorgievka, 4.X.1983 (A. Lelej); 1 ď, Novitzkoe, 22.VII.1984 (S. Belokobylskij); 1 Ψ, Nikolayevka, 7.VI.1986 (A. Lelej); 1 Ψ, Vysokogorsk, 12.VI.1986 (A. Lelej); 1 Ψ, Barabash-Levada, 8.VII.1986 (A. Lelej); 1 ď, Lazovskiy Reserve, 28.V–6.VI.2001 (M. Quest); 1 Ψ, the same place, 20.VII.2006 (P. Nemkov). Khabarovsk Terr.: 1 Ψ, Bureinskiy Mts., Suluk-Makit River, 7.VII.1988 (V. Mutin); 1 Ψ, Myaochan Mts., Amut Lake, 15.VII.1994 (V. Mutin). Amurskaya Prov.: 1 Ψ, Saskal, 13.VIII.1982 (A. Lelej). Sakhalin: 1 ď, Novikovo, 22.VII.1978 (A. Lelej); 1 Ψ, 9 km S Dolinsk, Sokol, 21.VIII.2003 (A. Lelej & S. Storozhenko) [ IBSS, ZIN].

Distribution. Russian Far East (Primorskiy Terr., Khabarovsk Terr., Amurskaya Prov., South Sakhalin). Etymology. Orientalis is a Latin adjective derived from the Orient, the East, referring to the area where the species has been found.

Figures

FIGURES 13 – 14. 13. Evagetes sahlbergi, Ψ, lectotype. 13 a: specimen and labels; 13 b: labels. 14. Evagetes orientalis sp. nov., ♂, holotype, specimen and labels.