Euryphlegon jacqueschassaini, , Otto, 2017

Otto, Robert L., 2017, A revision of Phlegoninae (Coleoptera: Eucnemidae), with descriptions of a new genus and four new species, Insecta Mundi 2017 (569), pp. 1-27 : 12-15

Euryphlegon jacqueschassaini sp. nov.

Fig. 33–37

Diagnosis. Apically straight pronotal hind angles distinguish E. jacqueschassaini from E. parallelus . Reddish-brown dorsum and Central American distribution further distinguishes it from E. degallieri . Type Material. Male holotype: “Barro Colorado I., CZ Panama, III, 13–18 1987, Hank Wolda” (month in Roman numeral and date range handwritten) / “Trap, 3B” / “ EUCNEMIDAE :, MACRAULACINAE :, Euryptychus , sp., Det. R.L. Otto, 2008” (year handwritten; folded) / “ HOLOTYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2016” (♂ handwritten behind species name on label) [red printed label]. Female allotype: “Barro Colorado I., CZ Panama, III, 23–25 1987, Hank Wolda” (month in Roman numeral and date range handwritten) / “Trap 4B” / “ ALLOTYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2016” (♀ handwritten behind species name on label) [yellow printed label]. Holotype and allotype are deposited in UCDIC.

Paratypes. 19 ♂♂, 16 ♀♀: NICARAGUA: 1 ♀, “Nica: Granada:, Volcan Mombacho, San Joaquin #1, 15-IV-98, Malaise trap col J.M. Maes” / “Collection of the Global, Eucnemid Research Project, (Robert L. Otto)” (green-framed white label) / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2016” (♀ handwritten behind species name on label) [yellow printed label] (GERP); PANAMA: 1 ♀, “Barro Colorado I., CZ Panama III, 7/10/12 1986, Hank Wolda” / “Trap, 4B” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2017” (♀ handwritten behind species name on label) [yellow printed label] (UCDIC); 2 ♂♂, “Barro Colorado I., CZ Panama III, 14/17/19 1986, Hank Wolda” / Trap, 4B” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2017” (♂ handwritten behind species name on label) [yellow printed label] (1, JMC; 1, UCDIC); 1 ♂, “Barro Colorado I., CZ Panama III, 14/17/19 1986, Hank Wolda” / Trap 2B” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2017” (♂ handwritten behind species name on label) [yellow printed label] (UCDIC); 3 ♀♀, “Barro Colorado I., CZ Panama III, 14/17/19 1986, Hank Wolda” / “Trap 4B” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2017” (♀ handwritten behind species name on label) [yellow printed label] (1, JMC; 2, UCDIC); 1 ♂, “Barro Colorado I., CZ Panama III, 21/24/26 1986, Hank Wolda” / “Trap 2A” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2017” (♂ handwritten behind species name on label) [yellow printed label] (UCDIC); 1 ♂, “Barro Colorado I., CZ Panama III, 21/24/26 1986, Hank Wolda” / “Trap 2B” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2017” (♂ handwritten behind species name on label) [yellow printed label] (UCDIC); 1 ♂, “Barro Colorado I., CZ Panama III, 21/24/26/3 1986, Hank Wolda” / “Trap 4B” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2017” (♂ handwritten behind species name on label) [yellow printed label] (UCDIC); 1 ♂, “Barro Colorado I., CZ Panama IV-2, III 28/31 1986, Hank Wolda” / “Trap 2B” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2017” (♂ handwritten behind species name on label) [yellow printed label] (UCDIC); 1 ♀, “Barro Colorado I., CZ Panama IV-2, III 28/31 1986, Hank Wolda” / “Trap 2B” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2017” (♀ handwritten behind species name on label) [yellow printed label] (UCDIC); 1 ♀, “Barro Colorado I., CZ Panama IV-2, III 28/31 1986, Hank Wolda” / Trap 4A” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2017” (♀ handwritten behind species name on label) [yellow printed label] (UCDIC); 1 ♂, “Barro Colorado I., CZ Panama III, 2–6 1987, Hank Wolda” (month in Roman numeral and date range handwritten) / “Trap, 4B” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2016” (♂ handwritten behind species name on label) [yellow printed label] (UCDIC); 1 ♂, “Barro Colorado I., CZ Panama III, 9–11 1987, Hank Wolda” (month in Roman numeral and date range handwritten) / “Trap, 4B” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2016” (♂ handwritten behind species name on label) [yellow printed label] (UCDIC); 2 ♂♂, “Barro Colorado I., CZ Panama III, 13–18 1987, Hank Wolda” (month in Roman numeral and date range handwritten) / “Trap, 2B” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2016” (♂ handwritten behind species name on each label) [yellow printed label] (UCDIC); 1 ♂, “Barro Colorado I., CZ Panama III, 13–18 1987, Hank Wolda” (month in Roman numeral and date range handwritten) / “Trap, 2B” / “Collection of the Global, Eucnemid Research Project, (Robert L. Otto)” (green-framed white label) / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2016” (♂ handwritten behind species name on label) [yellow printed label] (GERP); 1 ♀, “Barro Colorado I., CZ Panama III, 13–18 1987, Hank Wolda” (month in Roman numeral and date range handwritten) / “Trap, 4B” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2017” (♀ handwritten behind species name on label) [yellow printed label] (TAMU); 1 ♂ 1 ♀, “Barro Colorado I., CZ Panama III, 23–25 1987, Hank Wolda” (month in Roman numeral and date range and year handwritten) / “Trap, 2B” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2016” (either ♂ or ♀ handwritten behind species name on each label) [yellow printed label] (UCDIC); 1 ♂, “Barro Colorado I., CZ Panama III, 23–25 1987, Hank Wolda” (month in Roman numeral and date range handwritten) / “Trap, 4B” / “Collection of the Global, Eucnemid Research Project, (Robert L. Otto)” (green-framed white label) / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2016” (♂ handwritten behind species name on label) [yellow printed label] (GERP); 1 ♂ 1 ♀, “Barro Colorado I., CZ Panama IV, 1–3 1987, Hank Wolda” (month in Roman numeral and date range handwritten) / “Trap, 4A” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2016” (either ♂ or ♀ handwritten behind species name on each label) [yellow printed label] (UCDIC); 2 ♂♂, “Barro Colorado I., CZ Panama IV, 1–3 1987, Hank Wolda” (month in Roman numeral and date range handwritten) / “Trap, 2B” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2016 (or 2017)” (♂ handwritten behind species name on each label) [yellow printed label] (1, UCDIC; 1, TAMU); 1 ♀, “Barro Colorado I., CZ Panama IV, 1–3 1987, Hank Wolda” (month in Roman numeral and date range handwritten) / “Trap, 3B” / “Collection of the Global, Eucnemid Research Project, (Robert L. Otto)” (green-framed white label) / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2016” (♀ handwritten behind species name on label) [yellow printed label] (GERP); 1 ♀, “Barro Colorado I., CZ Panama IV, 6–10 1987, Hank Wolda” (month in Roman numeral and date range handwritten) / “Trap, 4A” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2016” (♀ handwritten behind species name on label) [yellow printed label] (UCDIC); 1 ♀, “Barro Colorado I., CZ Panama IV, 6–10 1987, Hank Wolda” (month in Roman numeral and date range handwritten) / “Trap, 2B” / “ Euryptychus , sp., Det. R. Otto, 2008” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2017” (♀ handwritten behind species name on label) [yellow printed label] (TAMU); 1 ♂, “Barro Colorado I., CZ Panama IV, 6–10 1987, Hank Wolda” (month in Roman numeral and date range handwritten) / “Trap, 2B” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2016” (♂ handwritten behind species name on label) [yellow printed label] (UCDIC); 1 ♀, “Barro Colorado I., CZ Panama IV, 6–10 1987, Hank Wolda” (month in Roman numeral and date range handwritten) / “Trap, 3B” / “Collection of the Global, Eucnemid Research Project, (Robert L. Otto)” (green-framed white label) / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2016” (♀ handwritten behind species name on label) [yellow printed label] (GERP); 1 ♂ 1 ♀, “Barro Colorado I., CZ Panama IV, 13–17 1987, Hank Wolda” (month in Roman numeral and date range handwritten) / “Trap, 2B” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2016” (either ♂ or ♀ handwritten behind species name on each label) [yellow printed label] (UCDIC); 1 ♀, “Barro Colorado I., CZ Panama V, 4–6 1987, Hank Wolda” (month in Roman numeral and date range handwritten) / “Trap, 2B” / “ PARATYPE:, Euryphlegon , jacqueschassaini, Otto , det. R.L. Otto, 2016” (♀ handwritten behind species name on label) [yellow printed label] (UCDIC). Paratypes are deposited in GERP, JMC, TAMU and UCDIC.

Description. Male holotype: Length 8.0 mm, width, 2.0 mm. Body oblong, elongate; uniformly dark reddish brown; antennae and legs dark reddish brown; head, pronotum and elytra clothed with short, recumbent yellowish setae ( Fig. 33 ).

Head: Subspherical, with median carina at vertex extending down to frons; surfaces shiny, punctures enlarged, evenly spaced, except at frontoclypeal region; apical margin of frontoclypeal region rounded, more than 2 times wider than base; mandibles stout, bidentate, densely punctate.

Antenna ( Fig. 34 ): Antennomeres IX–XI capitate, about 3/4 of body length; antennomere III longer than wide, slightly longer than II; antennomeres IV–V slightly longer than wide; antennomeres VI–VIII subequal, quadrate; antennomeres IX–XI each much longer than wide, each as long as combined length of V–VIII; lateral keel present on antennomere IX; antennomeres IX and X subequal; antennomere XI slightly longer than either IX or X, asymmetrical.

Pronotum: Surfaces shiny; punctures closely spaced, slightly rugose laterally; as long as wide, with large, sharp, apically straight hind angles; lateral sides gradually narrowing anteriorly, slightly constricted above hind angles; disc convex; base sinuous.

Scutellum: Short, setose, shallowly punctate, subtriangular and distally truncate to bilobed.

Elytra: Striae indicated as smooth lines at humeri and near apices, elsewhere punctate, somewhat obscured in middle; interstices flattened; surfaces shiny, transversely rugose; apices ( Fig. 35 ) with 15 deep, round cavities.

Legs: First tarsomere as long as combined lengths of remaining four on meso- and metatarsi; tibiae rounded in cross section; metatarsomeres I–III simple; metatarsomere IV excavated, as wide as III; metatarsomere V elongate; pretarsal claws simple.

Venter: Closely punctate, with elongate, recumbent yellowish setae; hypomeron simple, without lateral antennal grooves; metaepisterna parallel-sided; metacoxal plates medially 3.0–6.0 times wider than laterally.

Female allotype ( Fig. 36 ): 8.5 mm long, 2.5 mm wide; antennae ( Fig. 37 ) capitate, about 2/3 of body length; antennomere III longer than wide, slightly longer than II; antennomeres IV–V slightly longer than wide; antennomeres VI–VIII subequal, quadrate; antennomeres IX–XI each much longer than wide; lateral keel present on antennomeres IX and X; antennomere IX slightly longer than X, nearly as long as combined length of V–VIII; antennomere X as long as combined length of VI–VIII; antennomere XI distinctly shorter than X.

Variation. Male paratypes are 5.0–9.0 mm long and 1.5–3.0 mm wide, some are shorter and narrower than the holotype. Others are either subequal or larger and slightly wider than the holotype. Female paratypes are 6.0– 9.5 mm long and 2.0–3.0 mm wide, with some being shorter than and just as wide as or wider than the holotype. At least several females are larger and wider than the holotype. One female is as long as the holotype, but slightly wider. A small number of paratypes examined exhibit a weak median carina extending from the vertex to the frons. Most of the paratypes exhibit a strong median carina. Lateral keels on antennomeres IX–X exhibit some degree of variability. At least two paratypes lack any lateral keel on both antennomeres. Some have weakly indicated lateral keels on antennomeres IX and X. Most of the paratypes examined exhibit strong lateral keels on both antennal segments.

Distribution. This species is known from several trap localities on the Barro Colorado Island within the Canal Zone of Panama. One specimen was also taken in Nicaragua.

Biology. All specimens were taken from flight intercept traps, including a Malaise trap in Nicaragua. Larvae and pupae are unknown.

Etymology. The specific epithet is dedicated to my friend and colleague Jacques Chassain of Comesla-Ville, France, for his efforts in assisting me with answers to my many questions, and for providing specimens for my research.

Figures

Figures 33–37. Euryphlegon jacqueschassaini sp. nov. 33) Male holotype (UCDIC), dorsal habitus. 34) Male holotype antenna (UCDIC), lateral view. 35) Male holotype elytral apices (UCDIC), dorsal view. 36) Female allotype (UCDIC), dorsal habitus. 37) Female allotype antenna (UCDIC), lateral view. (Scale: 33–34, 36–37 = 1.0 mm; 35 = 5.0 mm)