Landouria chloritoides, Nahok & Tumpeesuwan & Tumpeesuwan, 2021
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publication ID |
https://doi.org/ 10.5852/ejt.2021.767.1495 |
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publication LSID |
lsid:zoobank.org:pub:9F019E02-95E7-4B6A-B5BC-01F8E26F3466 |
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DOI |
https://doi.org/10.5281/zenodo.5528094 |
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persistent identifier |
https://treatment.plazi.org/id/740CA614-D17B-47A3-A719-7DFC7D2CE2D5 |
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taxon LSID |
lsid:zoobank.org:act:740CA614-D17B-47A3-A719-7DFC7D2CE2D5 |
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treatment provided by |
Felipe |
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scientific name |
Landouria chloritoides |
| status |
sp. nov. |
Landouria chloritoides View in CoL sp. nov.
urn:lsid:zoobank.org:act:740CA614-D17B-47A3-A719-7DFC7D2CE2D5
Figs 3D View Fig , 4D View Fig , 5D View Fig , 6J–L View Fig ; Tables 1–4 View Table 1 View Table 2 View Table 3 View Table 4
Landouria sp.1 – Ounchareon 2015: 29–32, 44–50, 54–56, 59, 61, figs 4.2c, 4.3c, 4.4. 4.5i–l, 5.5, 5.7.
Landouria sp.2 – Buddharaksa 2016: 37–38, 41–42, figs 4.19, 4.20, 5.1, 5.2.
Landouria sp.4 – Yingkhamhang 2016: 25–26, 39–41, 43–45, 49–52, figs 4.7, 4.8, 4.21d, 4.22d, 5.3a, 5.4a.
Diagnosis
Shell shape similar to that of the genus Chloritis Beck, 1837 (Camaenidae) , but with last whorl stout and usually slightly angular, peripheral keel blunt, suture deep, aperture oval and oblique. Flagellum small ovate, penis very large and stout compared to vagina; vagina and free oviduct short ( Fig. 5D View Fig ). Radula with lanceolate teeth.
Etymology
The specific epithet ‘ chloritoides ’ refers to the conchological similarity between the shell of the new species and that of the camaenid genus Chloritis .
Type material
Holotype THAILAND • shell (SH = 12.0 mm, SW = 19.0 mm, AH = 7.7 mm, AW = 7.3 mm, UW = 3.9 mm); Khon Kaen Province, Chum Phae District, Wat Thep Udom Wanaram ; 16°45′29″ N, 101°57′55.03″ E; alt. 406 m; 9 Jun. 2017; B. Nahok, S. Tumpeesuwan and C. Tumpeesuwan leg.; NHMSU-00029 . ( Fig. 3D View Fig ) GoogleMaps
Paratypes THAILAND • 8 shells, 14 living specs; same collection data as for holotype; GenBank No. MN449404 View Materials and MZ435749 View Materials ; NHMSU-00030 GoogleMaps .
Other material examined
THAILAND • 4 shells, 8 living specs; North East, Khon Kaen Province, Chum Phae District, Pu Lup Cave ; 16°49′49.78″ N, 101°59′9.97″ E; alt. 336 m; NHMSU-00031 GoogleMaps .
Description
SHELL ( Fig. 3D View Fig , Table 1 View Table 1 ). Dextral, somewhat depressed, relatively large-sized. Whorls 6½, suture deep, apex obtuse, and moderately depressed, with low conical spire. Body whorl rounded, slightly keeled, red-brown in living specimens ( Fig. 4D View Fig ), empty shells are paler ( Fig. 3D View Fig ). Umbilicus very deep and wide. Aperture almost circular, with slightly thickened and weakly expanded lip.
GENITAL SYSTEM (n = 3) ( Fig. 5D View Fig , Table 4 View Table 4 ). Penis very large and stout, longer than vagina, cylindrical, gradually tapering towards epiphallus, internally with six rather thick, corrugated longitudinal pilasters; opening of the short, grooved verge surrounded by 2–3 lobes. Epiphallus short and thick. Flagellum short, elongate ovate, without nodes, internally with three large longitudinal pilasters. Vas deferens long and slender, entering in the lower half of the flagellum. Vagina short, as long as free oviduct, internally with seven, corrugated longitudinal pilasteres varying in size. Gametolytic sac thickened at base, with a long narrow cylindrical tube, and a small oval sac at distal end. Uterus swollen, prostate gland long.
RADULA (n = 3). Comprises 125–129 transverse rows with 75–83 teeth per row; radular formula: (26–28)- (11–13)-1-(11–13)-(26–28). Central tooth short lanceolate, symmetric, unicuspid ( Fig. 6J View Fig ). Lateral teeth large and elongated lanceolate ( Fig. 6J View Fig ). Marginal teeth lanceolate, gradually changing from unicuspid to bicuspid and finally tricuspid; endocone small; ectocone slightly oblique, with wide triangular notch and two to four lateral cusps ( Fig. 6K–L View Fig ).
Remarks
Landouria chloritoides sp. nov. can be distinguished from other species of Landouria in Thailand by its conchological similarity to the genus Chloritis and its much larger and stout penis ( Fig. 5D View Fig ). The radula of L. chloritoides sp. nov. is similar to that of L. tuberculata sp. nov. but differs by its unicuspid lateral teeth ( Fig. 5J View Fig ) vs the bicuspid lateral teeth of L. tuberculata sp. nov.
Distribution
Landouria chloritoides sp. nov. is currently known only from the limestone area around Phu Pha Man National Park, Chum Phae District, Khon Kaen Province, Thailand.
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Genus |
Landouria chloritoides
| Nahok, Benchawan, Tumpeesuwan, Sakboworn & Tumpeesuwan, Chanidaporn 2021 |
Landouria sp.2
| Buddharaksa W. 2016: 37 |
Landouria sp.4
| Yingkhamhang A. 2016: 25 |
Landouria sp.1
| Ounchareon J. 2015: 29 |