Metagonia maldonado, HUBER, 2000

HUBER, BERNHARD A., 2000, New World Pholcid Spiders (Araneae: Pholcidae): A Revision At Generic Level, Bulletin of the American Museum of Natural History 2000 (254), pp. 1-348 : 64-67

Metagonia maldonado , new species Figures 2 , 62 , 107 , 114 , 146 , 244 248

TYPES: Male holotype, 93 9♀ paratypes from 15 km E Puerto Maldonado (12°33'S, 69°03'W), Madre de Dios, Peru ; 200 m elev., June July 1989 and Feb. 25 Mar. 7, 1990 (D. Silva), in MUSM .

ETYMOLOGY: Named for the type locality. The specific name is a noun in apposition.

DIAGNOSIS: Representative of group 3 above. Easily distinguished from close relatives ( M. tingo , M. samiria ) by the tip of the procursus with its four pointed laminae and apophyses (figs. 244 245), by the shape and length of the femur apophysis (fig. 246), by the more widely separated clypeus apophyses (fig. 2), and by the shape of the epigynum (fig. 247).

MALE (holotype): Total length 3.2, carapace width 1.0; leg 1: 33.6 (8.3+0.4+8.1+15.2 +1.7), tibia 2: 4.8, tibia 3: 2.7, tibia 4: 4.5; tibia 1 l/d: 94. Habitus and prosoma shape as in M. tingo (cf. figs. 226 228), only posterior third of carapace brown, ocular area brown, distance PME-ALE about 20% of PME diameter. Clypeus brown, with pair of apophyses as in fig. 2 (see also fig. 107); sternum whitish. Chelicerae ochre-yellow, unmodified except pair of darker (sclerotized?) areas un- der clypeal apophyses. Palps ochre-yellow to light brown, procursus with dark distal parts; coxa without apophysis, femur with long prolateroventral apophysis and series of smaller apophyses on ventrodistal margin (fig. 246); procursus relatively simple (figs. 244 245), without hinged process; bulb simple, as typical for genus (fig. 245); tarsal organ capsulate (fig. 62). Legs ochre-yellow, with brown patellae and tibia-metatarsus joints; without spines, without curved and vertical hairs; retrolateral trichobothrium of tibia 1 at 6.9%; tarsus 1 with ~ 25 pseudosegments. Opisthosoma ochre-gray with some indistinct darker spots dorsally, shape as in M. tingo (cf. fig. 226). Male gonopore with four epiandrous spigots (fig. 114), ALS with several piriform gland spigots (fig. 146).

VARIATION: The males from Tambopata, Beni, and La Paz (see below) differ slightly with respect to the dorsal spine of the procursus tip (arrow in fig. 244), which is slight- ly narrower, and with respect to the series of ventrodistal apophyses on the femur (most conspicuously, there is an additional upwardpointing apophysis where the arrow points to in fig. 246).

FEMALE (paratypes): Tibia 1 (N = 8) 6.0 7.1 (x¯ = 6.5). In general very similar to male, but carapace and clypeus without distinct brown marks, only light brown spot posteriorly and light brown band between eye triads. Epigynum light brown with dis- tinct darker marks and lateral margins (fig. 247). Internally apparently symmetric, with two pairs of folds that seem to originate from uterus externus (fig. 248).

DISTRIBUTION: Known from southern Peru (Madre de Dios) and northern Bolivia (Beni, La Paz) (map 1).

MATERIAL EXAMINED: PERU: Madre de Dios: 15 km E Puerto Maldonado: types above. Zona Reservada Tambopata, Laguna Chica : noche (12°50'S, 69°17'W), 290 m elev., June 8, 1988 (D. Silva), 13 1♀ in MUSM ; Zona Reservada Tambopata, June 3 15, 1988 (3 vials; J. Coddington), 13 2♀ in USNM ; same locality, Nov. 7 12, 1983, and Sept. 6 14, 1984 (3 vials; T. L. Erwin et al.), 13 2♀ in USNM ; Tambopata Reserve, Rio Tambopata, Explorer s Inn , Mar. 1988 (J. Palmar, D. Smith), 13 in MCZ ; Zona Reservada de Manú, Rio La Torre and Rio Tambopata (12°50'S, 69°17'W), Aug .

Dec. 1979 (A. Rypstra), 6♀ in USNM. BO- LIVIA: Beni: 16.8 mi SW Yucumo (~ 15°23'S, 66°59'W), ~ 500 m elev., Nov. 15 19, 1989 (2 vials; J. Coddington, C. Griswold, D. Silva, S. Larcher, E. Penaranda), 53 7♀ in USNM ; La Paz: Cerro Uchumachi , 7 km SW Coroico, (~ 16°15'S, 67°21'W), ~ 1900 m elev., Nov. 24 25, 1989 (J. Coddington, C. Griswold, D. Silva, S. Larcher, E. Penaranda), 33 12♀ in USNM .

Figures

Figs. 1–6. Carapace shape and eye pattern, frontal view. 1. Metagonia delicata (O. Pickard-Cambridge), female. 2. Metagonia maldonado, n. sp., male. 3. Papiamenta levii (Gertsch), female. 4. Ninetis namibiae, n. sp., female. 5. Holocnemus hispanicus Wiehle, female. 6. Tupigea lisei, n. gen., n. sp., female. Scale lines: 0.3 mm.

Figs. 61–66. Tarsal organs in Metagonia (all on male pedipalp). 61. M. blanda Gertsch. 62. M. maldonado, n. sp. 63. M. delicata (O. Pickard-Cambridge). 64. M. tinaja Gertsch. 65. M. globulosa, n. sp. 66. M. argentinensis Mello-Leitão. Scale lines: 15 µm.

Figs. 105–112. Miscellaneous structures. 105. Leptopholcus dalei (Petrunkevitch), modified hair at tip of male palpal trochanter apophysis. 106. Micropholcus fauroti (Simon), modified hair at tip of male palpal trochanter apophysis. 107. Metagonia maldonado, n. sp., tip of clypeal apophysis. 108. Modisimus dominical Huber, male femur, showing vertical (and ‘‘normal’’) hairs. 109. Ibotyporanga naideae Mello- Leitão, male tibia, showing vertical (and ‘‘normal’’) hairs. 110. Coryssocnemis simla, n. sp., spine on male femur 2. 111–112. Modisimus dominical Huber, pseudotrichia on inner surface of male palpal coxae (endites). Scale lines: 100 µm (108–109), 20 µm (107, 110), 5 µm (105–106, 111–112).

Figs. 113–118. Male gonopore with epiandrous spigots in Metagonia. 113. M. argentinensis Mello- Leitão. 114. M. maldonado, n. sp. 115. M. delicata (O. Pickard-Cambridge). 116. M. blanda Gertsch. 117. M. uvita Huber. 118. Metagonia rica Gertsch. Scale lines: 30 µm (113–114, 116–118), 10 µm (115).

Figs. 146–151. Anterior lateral spinnerets in Metagonia spp., and in Chisosa diluta. 146. M. maldonado, n. sp., male. 147. M. argentinensis Mello-Leitão, male. 148. M. blanda Gertsch, male. 149. M. rica Gertsch, male. 150. M. globulosa, n. sp., female. 151. Chisosa diluta (Gertsch and Mulaik), female. Scale lines: 15 µm.

Figs. 244 248. Metagonia maldonado, n. sp. 244. Left procursus, retrolaterodorsal view (arrow: this structure is slightly narrower in males from Uchumachi, Tambopata, and Yucumo). 245. Left male palp, retrolateral view. 246. Male left palpal femur, retrolateral view (arrow: at this point, males from Uchumachi, Tambopata, and Yucumo have a small, additional apophysis). 247. Epigynum, ventral view. 248. Epigynum, dorsal view. Scale lines: 0.2 mm.