Anaspis curva, Levey, 2020

Levey, Brian, 2020, A review of the species of Anaspis (s. str.) similar to A. nigripes Brisout and A. apfelbecki Schilsky, with the description of three new species (Coleoptera: Scraptiidae), Zootaxa 4778 (3), pp. 509-520 : 514-515

Anaspis curva sp. n.

( Figs. 4 , 10 , 19 , 29 )

Holotype ♂ ( NMGW) China: Beijing, Dongling Mts., Xialongmen , Mei Yao Yu, N 39ᵒ 96′, E115ᵒ 43′. 16.vi.2001. J. Cooter. / On flowers of Hydrangea bretschniederi Dipp ./ Holotype Anaspis curva sp. n. B. Levey 2015. Paratypes 2♀ ( BMNH, NMGW) Beijing Region: Xialongmen. ca 1100m. N39 o 58.074′ E115 o 25.882′. 9–12.vi.2004. J. Cooter .

Diagnosis. General diagnosis of holotype (male) ( Fig. 29 ): Length (fore margin of pronotum to apex of elytra in mid line) 3.0 mm. Head from level of antennal insertions, scutellum, elytra and underside black, frons below antennal insertions, labrum, maxillary palpi except for distal segment, basal three segments of antennae, coxae, trochanters and basal two-thirds of fore legs and pronotum except for medially in basal two-thirds, yellow; pronotum medially in basal two-thirds broadly dark brown; distal seven segments of antennae black; distal segment of maxillary palpi, distal third of fore femora, tibia and tarsi dark brown; mid and hind legs entirely dark brown to black. Entire body densely covered with short grey pubescence.

Head. Antennae: first antennomere almost twice as long as wide, nearly rectangular; second antennomere cylindrical, slightly longer than first; third and fourth antennomeres about equal length, cylindrical, slightly widened at apex, about 1.5× as long as second; fifth and sixth antennomeres about equal length, cylindrical, slightly shorter and broader than fourth; seventh antennomere about equal to sixth, slightly triangular; eighth to tent antennomere becoming progressively shorter and broader, nearly triangular; eleventh antennomere ovate, about 1.6× as long as tenth. Apical segment of maxillary palp securiform. Pronotum: About 1.30× as wide at base as long in midline; hind angles almost right-angled; lateral edge straight when viewed from side. Scutellum: Triangular about as wide as long. Elytra: 3.8× as long as pronotum. Abdomen ( Fig. 19 ): Ventrites 1 and 2 convex; 3 triangular in cross section, with a slightly developed rounded median keel, with a large projecting tuft of laterally divergent setae in mid line and a pair of long, strongly curved, not twisted, appendages, widely separated at the base; 4 weakly convex, the posterior margin narrowly, moderately deeply emarginated medially; 5 deeply emarginated at centre. Legs: fore leg ( Fig. 4 ), with femora very broad and strongly flattened anterior-posterior and slightly excavated on the posterior surface; tibia about three-quarters length of femora, slightly flattened anterior-posterior, slightly widening distally with a dense distal fringe of dark brown spines and two yellow spurs the longest slightly longer than tarsal segment 1; tarsal segment 1 small, about as wide as long, slightly widening distally; 2 about 1.5× as long as 1, as wide at base as 1 at apex, widening distally, about as wide as long, with a ventral pad extending distally beyond the base of the segment 3; 3 about as long as 2, approximately heart shaped with a deep medial excision distally, about 1.25× as wide at apex as long, with a long semicircular ventral pad extending distally beneath the entire length of segment 4; 4 very small oblong, slightly longer than wide; 5 elongate, club shaped about as long as 3 and 4 together, claws small curved, slightly widened at base. Mid leg with femora moderately anterior-posterior flattened; tibia about as long as femora, regularly widening from base to apex, the ventral margin almost straight, slightly sinuate near the apex; with a dense distal fringe of yellow-brown spines and a pair of long yellow spurs, the longer about half the length of tarsal segment 1, the shorter about one-third length of segment 1; tarsal segment 1 long, cylindrical, 0.53 length of tibia; 2 shorter, cylindrical, 0.56 length of 1; 3 shorter, 0.50 length of 2, approximately heart shaped with a deep medial excision, with a long semicircular ventral pad; 4 very small oblong, slightly longer than wide; 5 elongate, club shaped about as long as 3 and 4 together, claws small curved, slightly widened at base. Hind leg with femora strongly flattened anterior-posterior; tibia about half three quarters length of femora, strongly widened from base to apex, about 4.0× as long as wide at apex, with a dense distal fringe of short brown spines and two yellow spurs, the longer about two-thirds the length and the shorter about half the length of tarsal segment 1; tarsal segment 1 moderately widening at apex, about four× as long as wide at apex, about 0.70 length of tibia; 2 slightly widening at apex, about 0.6 length of 1; 3 very slightly widening at apex, about 0.60 length of 2; 4 cylindrical, about as long as 3, claws small, slightly curved, slightly widened at base. Genitalia ( Fig. 10 ): Parameres hooked at apex; basal piece 1.2× as long as parameres.

General diagnosis of female: Length 2.8–4.1 mm. Colour as in male; smaller female with pronotum entirely yellow. Head: same as male but seventh to tenth antennomeres slightly shorter and broader. Pronotum: same as male. Scutellum: same as male. Elytra: 3.5–3.7× length of pronotum. Abdomen: ventrites 1–4 convex, not keeled; ventrite 5 approximately triangular with a rounded apex. Legs: fore leg with femora strongly flattened anteriorposterior; tibia about three-quarters length of femora, slightly flattened anterior-posterior, slightly widening distally with a dense distal fringe of yellow-brown spines and two yellow spurs the longest slightly longer than half length of tarsal segment 1; tarsal segment 1 elongate, slightly widening distally, about 3.0× as long as wide; 2 about twothirds length of 1, and slightly wider; 3 about as long as 2, approximately heart shaped with a deep medial excision distally, about as wide at apex as long, with a long semicircular ventral pad extending distally beneath the entire length of segment 4; 4 very small oblong, slightly longer than wide; 5 elongate, club shaped about as long as 3 and 4 together, claws small curved, widened at base. Mid leg as in male but tibial spurs and tarsi slightly shorter relative to the length of the tibia. Hind leg with femora strongly flattened anterior-posterior; tibia about half length of femora, strongly widened from base to apex, about 2.5× as long as wide at apex, with a dense distal fringe of short yellow spines and two yellow spurs, the longer about two-thirds the length and the shorter about half the length of tarsal segment 1; tarsal segment 1 moderately widening at apex, about three to four× as long as wide at apex, about 0.70 length of tibia; 2 slightly widening at apex, about 0.70 length of 1; 3 very slightly widening at apex, about 0.70 length of 2; 4 cylindrical, about as long as 3, claws small, slightly curved, slightly widened at base.

Comparative diagnosis. Closely related to A. bertrami sp. n. (see comparative diagnosis under this species).

Etymology. This species is named for the more strongly curved appendages of ventrite 3 in the male, in comparison to those of A. bertrami sp. n.

Figures

FIGURES 1–9. Fore legs of male: 1. A. emarginata Schilsky, N. China, Heilunkiang, Erlungshan (BMNH); 2. A. nigripes Brisout de Barneville, France, Côte-d’Or, Pouilly –en-Auxois (NMGW); 3. A. cooteri sp. n., holotype (NMGW); 4. A. curva sp. n., holotype (NMGW); 5. A. bertrami sp. n., holotype (NMGW); 6. A. graeca Schilsky, syntype (BMNH); 7. A. apfelbecki Schilsky, paralectotype (ZMHB); 8. A. apfelbecki Schilsky, paralectotype (ZMHB); 9. A. marseuli Csiki, Japan, Tochigi, Tsurutanuma, Utsunomiya City (TPMJ).

FIGURES 10–17. Male genitalia: 10. A. curva sp. n., holotype (NMGW); 11. A. bertrami sp. n., holotype (NMGW); 12. A. apfelbecki Schilsky, paralectotype (ZMHB); 13. A. apfelbecki Schilsky, paralectotype (ZMHB); 14. A. cooteri sp. n. holotype (NMGW); 15. A. nigripes Brisout de Barneville, France, Côte-d’Or, Pouilly-en-Auxois (NMGW); 16. A. koenigi Schilsky, paralectotype (ZMHB); 17. A. marseuli Csiki, Japan, Tochigi, Yunishikawa, Kuriyama Vill. (TPMJ).

FIGURES 18–27. Apical abdominal ventrites of male: 18. A. apfelbecki Schilsky, paralectotype (ZMHB); 19. A. curva sp. n., holotype; 20. A. bertrami sp. n., holotype (NMGW); 21. A. emarginata Schilsky, N. China, Heilunkiang, Erlungshan (BMNH); 22. A. nigripes Brisout de Barneville, Spain: La Rioja, Nieva de Cameros, La Dehesa (NMGW); 23. A. koenigi Schilsky, paralectotype (ZMHB); 24. A. marseuli Csiki, Japan, Tochigi City (TPMJ); 25. A. marseuli Csiki, Japan, Utsunomiya City (TPMJ); 26. A. cooteri sp. n., paratype (NMGW); 27. A. graeca Schilsky, syntype (BMNH).

FIGURES 28–36. Dorsal habitus: 28. Anaspis bertrami sp. n. female paratype (NMGW); 29. A. curva sp. n., male holotype (NMGW); 30. A. emarginata Schilsky, N. China, Heilunkiang, Erlungshan (BMNH); 31. A. cooteri sp. n., male paratype (NMGW); 32. A. marseuli Csiki, Japan, Hyogo, Haruki, Onsen Town (TPMJ); 33. A. nigripes Brisout de Barneville, male, France, Côte-d’Or, Pouilly-en-Auxois (NMGW); 34. A. apfelbecki Schilsky, paralectotype (ZMHB); 35. A. koenigi Schilsky, paralectotype (ZMHB); 36. A. graeca Schilsky, syntype (BMNH).