Sciarokeroplatus L. PAPP et ŠEVČÍK, 2005

Papp, L. & Ševčík, J., 2005, Sciarokeroplatinae, A New Subfamily Of Keroplatidae (Diptera) *, Acta Zoologica Academiae Scientiarum Hungaricae 51 (2), pp. 113-123 : 113-116

Sciarokeroplatus L. PAPP et ŠEVČÍK , gen. n.

Type species: Sciarokeroplatus pileatus L. PAPP et ŠEVČÍK , sp. n.

Gender: masculine.

The terminology for the structure of the head follows that used for Macrocera by MATILE (1990), but our use of this terminology should not be regarded as a definite acceptance of homologies for the head parts of Macrocera . Frons (postfrons) short, cerebral sclerite large, occupying all dorsal part of head. There is a definite furrow (suture) between postfrons and cerebral sclerite (? plus occiput) ( Fig. 1 ). This “vertical” suture is placed rather anteriorly: it runs below dorsal margin of eyes. Consequently, head seems “capped”, this is why the specific epithet of the type species is pileatus . No ocelli. No eye-bridge above the antennae. A pair of small shallow emarginations are discernible above scape (more on holotype, less seen on paratype), whose function is unknown but they definitely do not correspond to the ocelli. Postgena (? occiput + postgena) large, bulging, distance of left and right halves only 0.04 mm above foramen. Foramen small.

Face extremely short, similar to that of Macrocera spp. Clypeus ( Fig. 2 ) largely pentagonal, high with some medium-long setae.

Antenna with 14 flagellomeres ( Figs 1–2 ), scape and pedicel more or less cuboid. Flagellomeres simple.

Mouthparts very small, labella small, fleshy. Palpus four-segmented ( Fig. 2 ): basal segment short, second segment larger and thick, third and fourth segments long and thin cylindrical, ventrally with stumpy sensilla.

Wing membrane microtrichia even but wholly irregular ( Figs 3–4 ). No strong costal fringe. Costa thick, reaches 1/3 of distance between R 5 and M1 (wing vein terminology follows KRZEMIŃSKI & EVENHUIS 2000). Radial veins dorsally with long setae. Vein R 4 present ( Fig. 3 ). Cross-vein M–Cu, connecting common trunk of Rs and M1+2 stem vein with base of common trunk of M3 and Cu1, discernible only as a colourless fold ( Fig. 4 ). Base of M2 faint. M1–M2 fork apparently emerges from Rs (due to presence of a radiomedial fusion in which crossvein R –M is obliterated), M3 emerges from Cu1. Cu2 distinct ( Figs 3–4 ) and runs rather far from Cu1. A1 long but also A2 discernible to about 4/5 distance to anal margin. Base and insertion of the two anal veins are completely separated. A distinct alula present between anal angle and wing scale. Alula bears long setulae.

Stalk of halter medially with a subdorsal peg. Knob large, globose.

Femoral and tibial setulae simple, fine and rather long, and not ordered into rows. Coxae remarkably short, the mid and hind coxae about two-thirds as long as the fore coxa. The length of hind coxa about two-thirds of the height of the laterotergite. Tibial spurs 1+2+2, well developed and hairy. Fore tibia anteroapically with a large depressed or evenly emarginated area ( Fig. 5 ) with numerous small pointed setae, similar to Sciaridae , but much more numerous and not ordered into rows as in Sciaridae or in other families of Sciaroidea.

No anterior or posterior combs of small thornlets on mid and hind tibiae. Tarsal claws simple, normal, basal half almost straight. Well-developed pulvilli present; they are as hairy as empodium ( Fig. 6 ).

Abdomen with 8 normal preabdominal segments. Both tergites and sternites broad, in each segment tergite and sternite almost meet laterally, i.e. intervening membranous area rather small. Spiracle pairs 1–7 are situated in this membrane. Tergite 8 transverse, only half as long as sternite 8 or tergite 7. Tergite 9 shorter than long, rounded apically. Sternite 9 very short ( Fig. 7 ).

clypeus. Scale: 0.5 mm for Fig. 1 , 0.2 mm for Fig. 2 .

Male genitalia extremely simple. Tergite 9 medium-sized, simple. Cerci large and vertical, fleshy with small setulae only ( Figs 9–10 ). Left and right gonocoxites not fused medially ( Fig. 7 ), gonocoxite each similar to a punt with medial seat, i.e. the two medial edges of gonocoxite connected by a concave sclerite. Gonocoxal apodemes not particularly long. Gonostylus similar to a thumb of a glove, with setae and medial apical apex with two flat sharp tooth-like projections ( Figs 8 , 12 ). Aedeagus ( Fig. 11 ) simple, comparatively rather small, mostly membranous, apex truncated, with short hairs. Parameres small, weakly sclerotized.

Figures

Figs 1–2. Sciarokeroplatus pileatus sp. n., type males. 1 = head of holotype sublaterally (as if left flagellum were removed, del. A. Szappanos); 2 = head of a paratype, front view, perpendicular to

Fig. 3. Sciarokeroplatus pileatus sp. n., paratype male. 3 = photo of left wing dorsally

Figs 4–6. Sciarokeroplatus pileatus sp.n., paratype male. 4 = basal portion of wing, in higher magnification, 5 = apex of fore tibia, inner (medial) view, 6 = apex of fore tarsus, ventral view. Scales: 0.4

Figs 7–9. Sciarokeroplatus pileatus sp. n., paratype male, genitalia. 7 = gonostylus in ventral view (Taiwanparatype), 8 = genitalia in dorsal view, 9 = ventral view.Scale:0.2 mm for Fig.9, 0.1 mm for

Figs 10–12. Sciarokeroplatus pileatus sp. n., paratype male, genitalia. 10 = anal parts, ventral view (as if apex of cerci cut off), 11 = aedeagus and parameres with gonocoxal apodemes, dorsal view (T9