Ancyropilus emmotti, Bartlett & Lambkin, 2022

Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, Zootaxa 5220 (1), pp. 1-81 : 13-14

Ancyropilus emmotti sp. nov.

ZooBank registration: urn:lsid:zoobank.org:act:18D73014-582C-423F-8E7C-F74742DE7F3E

( Figs 28 , 67 , 103 , 139 ; Map 1)

HOLOTYPE ♁: Queensland: Qld: 24.142° Sx 143.196°E, 4km SSE HS Noonbah Station (NB3 M). 188m, 19Jan-7Feb2009. Malaise. A. Emmott. 17255. Sandy plain, Ghost gums ( QM, type reg. T258545) . PARATYPES (33): Queensland:same data as holotype (5, QM) ; Qld: 3.7km S HS Noonbah Station (NB4 M), 186m, 24.141° Sx 143.19°E, 19Jan-7Feb2009. Malaise. A. Emmott. Dense Gidgyea scrub. 17254 (1 ♀, 1, QM) ; Qld: 26.728° Sx 142.651°E Plevna Downs , Tompilly Hill Base (PD6 M) 187m 24Apr-24Jul 2008. G Turner, R Mackenzie. Malaise. Gidgee. 17320 (1 ♀, 1, QM) ; Qld: 26.541° Sx 142.516°E Plevna Downs , 2.3km SSW Arima (PD2 M) Malaise. 25Nov-17Dec 2008. Gidgee. R. Mackenzie. 134m. 17292 (1, QM) ; Qld: 26.541° Sx 142.516°E Plevna Downs , 2.3km SSW Arima (PD2 M) Malaise. 13 Jan-3 Apr 2008. 132m. R.Mackenzie. Gidgee 17284 (1, QM) ; Qld: 26.67° Sx 142.577°E 2.5km WNW Plevna Downs HS (PD3 M) 133m. Gidgee 15 Mar-3 Apr 2009. Malaise. R. Mackenzie. 17317 (1, QM) ; Qld: 26°40.3′ Sx 142°34.6′E Plevna Downs Hstd, 2.5km WNW. Malaise Trap, Gidgee. 12-29 Dec 2007. 130m. N. Starick, Lambkin & R. Mackenzie. 15998 (1 ♀, QM) ; Aust : QLD: 40km N Noccundra, dry creek bed 27°29″56′S:142°38″24′E 02-09-i-99 Malaise trap P. Bouchard, C. Lambkin, D. Yeates, N. Power (1, QM) ; Australia: n Qld Mt. Isa (within 10km) 23-31.XII.1990 R.I. Storey at light (1, QDPC) . New South Wales: NSW: 29.047° Sx 146.996°E Culgoa NP, 8km WNW Cawwell HS, Diemunga Lagoon (CGN2M) Malaise. 22Nov-21Dec 2009 C.Lambkin, B.Schiebaan N.Starick. Coolibah 19302 (1, QM) . Northern Territory: 23.59 S 133.56 E N.T. Ewaninga Area M. V. Light 19 Dec. 1987 J. & I. Archibald (3, NTMAG) .

Diagnosis. Pronotum more angulate than rounded laterally, disc heavily punctate, glabrous areas in middle, at base and either side of central impression; elytra dark with orange fasciate and apical maculations (humeral maculae absent), fascia convergent towards, and meeting at, the suture, punctation lacking nodules, 8 th stria beginning within fascia, all striae reaching apical maculae; femora yellow basally, brown apically, tarsi with three ventral tarsal pads.

Description. Habitus: Fig. 139 . Total length: 6.7–9 mm (holotype 7 mm). Head: Cranium black, submentum and gula often orange to orange-brown, clypeus and supra-antennal elevation with a deep reddish hue, anteclypeus and labrum orange, palpi and antennae orange-brown to brown; eyes separated by 0.36–0.6 eye widths (holotype 0.5); vertex and frons densely punctate, between eyes punctate-rugulose, clypeus with few punctations; genae and submentum wrinkled; ratio of exterior to interior edges of terminal palpomeres about 1.7:1 (maxillae) and 2:1 (labium); antennae not reaching base of pronotum; eyes and most of cranium vested with erect pale setae, frons with shorter finer medially-directed setae. Prothorax: Blackish to dark-brown; pronotum 1.14–1.26 times longer than wide (holotype 1.16), sides weakly angulate, middle wider than pronotal arch; subapical depression v-shaped, meeting shallow central impression; disc densely punctate, a medial glabrous stripe from base to central impression, area either side of central impression weakly tumescent, partially glabrous; short fine multi-directional setae (forming subtle swirls adjacent to glabrous areas) and long erect setae. Pterothorax: Ventrites brown to orange, with numerous pale, posteriorly- to posteromedially-directed setae; elytra blackish-brown with orange markings (each elytron with a large apical macula and an angulate transverse fascia which narrows towards, and meets at, the suture—its anterior margin at basal one-third); length to width ratio 2.43–2.67:1 (holotype 2.66); 8 th stria beginning within fascia, between seventh and eleventh punctation of 7 th stria, all striae ending at or within apical macula, punctation lacking nodules, punctations posterior of fascia slightly smaller than those anterior to fascia, epipleurae extending into apical maculae, interstriae with very fine short semi-erect setae and less frequent longer thicker erect setae, several setae often across interstical width; hindwing with CuA 3+4 and CuA 1 cross-veins complete, MP 3+4 basad of CuA 1 crossvein small (but not completely absent). Legs: Basal two-thirds of femora yellow, apical third dark brown (on average); tibiae and tarsi dark brown, ventral tarsal pads yellowish; meso- and metafemora slender, profemora slightly swollen. Abdomen: Ventrites orange. Male genitalia: Tegmen ( Fig. 28 ) broadest near middle, narrowing before expanding again before apically acuminate parameroid lobes, dorsal sinus about one-fifth tegmen length, wide open, sides slightly convergent, internal lateral opening with short membranous lobes bearing a brushlike row of very short setae, ventral sinus deeper (about one-quarter tegmen length), length of apodeme unclear due to gradual convergence of tegminal arms (though seemingly a little less than one-quarter tegmen length); median lobe as in Fig. 67 ; pygidium as in Fig. 103 .

Variation. The punctures of the 7 th stria, within the area anterior of the elytral fascia, in one specimen from Noonbah, are not linear but are staggered in a ‘zig-zag’ arrangement. This may incorrectly be interpreted as representing a broken 8 th stria, however, on this specimen the 8 th stria clearly begins just posterior of the anterior margin of the fascia.

Etymology. We name this species after Mr Angus Emmott, avid naturalist, and owner of Noonbah Station near Longreach.

Biology. Specimens were collected in October and November (NSW), November to July (Qld), November and December (NT), August, November and February (WA).

Distribution. Map 1 shows the distribution of Ancyropilus emmotti sp. nov. type specimens only. Additional specimens, from New South Wales, the Northern Territory and northern Western Australia were examined, but not designated as types.

Remarks. The species group name Notopilo emmotti , published in the 2009-2010 Bush Blitz survey report for north-western New South Wales and southern Queensland ( ABRS 2014a) with no accompanying description and without reference to a name-bearing type, is deemed unavailable ( ICZN 1999, Article 16.4).

Figures

FIGURES 27–47. Tegmina, dorsal: (27) Ancyropilus brigalowae, paratype; (28) Ancyropilus emmotti, holotype; (29) Ancyropilus exossuarius, paratype; (30) Ancyropilus monteithi, holotype; (31) Ancyropilus noonbahensis, holotype; (32) Ancyropilus packsaddlensis, holotype; (33) Ancyropilus simplex, paratype; (34) Ancyropilus tricolor, paratype; (35) Ancyropilus hornensis, paratype; (36) Culcipternotus mareebensis, paratype; (37) Infectostriatus absentis, holotype; (38) Infectostriatus differens, holotype; (39) Monilonotum bunyense, paratype; (40) Monilonotum doddi; (41) Monilonotum pascoei; (42) Monilonotum sundholmi, paratype; (43–47) Notopilo, beswickensis species group—(43) Notopilo beswickensis, holotype; (44) Notopilo calicis, paratype; (45) Notopilo tompricensis, holotype; (46) Notopilo xanthoimprocerus, holotype; (47) Notopilo xanthoprolatus, holotype (scale bars: 0.5 mm).

FIGURES 66–101. Median lobes: (66) Ancyropilus brigalowae, paratype; (67) Ancyropilus emmotti, holotype; (68) Ancyropilus exossuarius, paratype; (69) Ancyropilus monteithi, holotype; (70) Ancyropilus noonbahensis, holotype; (71) Ancyropilus packsaddlensis, holotype; (72) Ancyropilus simplex, paratype; (73) Ancyropilus tricolor, holotype; (74) Ancyropilus hornensis, paratype; (75) Culcipternotus mareebensis, paratype; (76) Infectostriatus absentis, holotype; (77) Infectostriatus differens, holotype; (78) Monilonotum bunyense, paratype; (79) Monilonotum pascoei; (80) Monilonotum sundholmi, paratype; (81–85) Notopilo, beswickensis species group—(81) Notopilo beswickensis, holotype; (82) Notopilo calicis, paratype; (83) Notopilo tompricensis, holotype; (84) Notopilo xanthoimprocerus, paratype; (85) Notopilo xanthoprolatus, holotype; (86–88) Notopilo, cambageicola species group—(86) Notopilo cambageicola, paratype; (87) Notopilo interfabulatus, paratype; (88) Notopilo tanybasilaris, paratype; (89) Notopilo congruus (congruus species group); (90) Notopilo reduncus (reduncus species group); (91–93) Notopilo, variipes species group—(91) Notopilo brevistriatus, paratype; (92) Notopilo gerstmeieri, holotype; (93) Notopilo variipes; (94–96) Notopilo, unplaced to species group—(94) Notopilo elstoni, holotype; (95) Notopilo katherinensis, holotype; (96) Notopilo obesus, paratype; (97) Platynotum bulli, paratype; (98) Platynotum culgoense, holotype; (99) Platynotum femorale; (100) Platynotum foveosetosa, holotype; (101) Platynotum gracile, holotype (scale bars: 0.5 mm).

FIGURES 102–137. Male pygidiae: (102) Ancyropilus brigalowae, paratype; (103) Ancyropilus emmotti, holotype; (104) Ancyropilus exossuarius, paratype; (105) Ancyropilus monteithi, holotype; (106) Ancyropilus noonbahensis, holotype; (107) Ancyropilus packsaddlensis, holotype; (108) Ancyropilus simplex, paratype; (109) Ancyropilus tricolor, paratype; (110) Ancyropilus hornensis, paratype; (111) Culcipternotus mareebensis, paratype; (112) Infectostriatus absentis, holotype; (113) Monilonotum doddi; (114) Monilonotum pascoei; (115) Monilonotum sundholmi, paratype; (116–120) Notopilo, beswickensis species group—(116) Notopilo beswickensis, holotype; (117) Notopilo calicis, paratype; (118) Notopilo tompricensis, holotype; (119) Notopilo xanthoimprocerus, paratype; (120) Notopilo xanthoprolatus, holotype; (121–124) Notopilo, cambageicola species group—(121) Notopilo cambageicola, paratype; (122) Notopilo interfabulatus, paratype; (123) Notopilo lawnhillensis, paratype; (124) Notopilo tanybasilaris, holotype; (125) Notopilo congruus (congruus species group); (126–128) Notopilo, variipes species group—(126) Notopilo brevistriatus, paratype; (127) Notopilo gerstmeieri, holotype; (128) Notopilo variipes; (129–132) Notopilo, unplaced to species group—(129) Notopilo confusus, holotype; (130) Notopilo elstoni, holotype; (131) Notopilo katherinensis, holotype; (132) Notopilo obesus, paratype; (133) Platynotum bulli, paratype; (134) Platynotum culgoense, holotype; (135) Platynotum femorale; (136) Platynotum foveosetosa, holotype; (137) Platynotum gracile, holotype (scale bars: 0.5 mm).

FIGURES 138–149. Habitus: (138) Ancyropilus brigalowae, paratype; (139) Ancyropilus emmotti, holotype; (140) Ancyropilus exossuarius, paratype; (141) Ancyropilus hornensis, paratype; (142) Ancyropilus labris, holotype; (143) Ancyropilus monteithi, paratype; (144) Ancyropilus noonbahensis, holotype; (145) Ancyropilus packsaddlensis, holotype; (146) Ancyropilus simplex, paratype; (147) Ancyropilus tricolor, paratype; (148) Culcipternotus mareebensis, holotype; (149) Infectostriatus absentis, holotype (scale bars: 1.0 mm).