Asmodeus petrasnerus Seoane and Cerdeño, 2014
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publication ID |
https://doi.org/ 10.4202/app.00301.2016 |
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persistent identifier |
https://treatment.plazi.org/id/03ADD838-FF80-FF8F-8B15-F9F1FE00873A |
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treatment provided by |
Felipe |
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scientific name |
Asmodeus petrasnerus Seoane and Cerdeño, 2014 |
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Asmodeus petrasnerus Seoane and Cerdeño, 2014
Fig. 7A View Fig .
Material.—MCNAM-PV4958,leftcalcaneum;MCNAM-PV 5077, fragment of left calcaneum from Quebrada Fiera, Mendoza, Argentina, Deseadan SALMA, late Oligocene.
Emended diagnosis.—The calcaneum MCNAM-PV 4958 allows establishing new characteristics for A. petrasnerus , which is reflected in the following emended diagnosis. Calcaneum with well-developed tendinous sulcus in the tuber and anterior border of the neck projected and with a distal semicircular notch, both features differing from Asmodeus osborni and Homalodotherium ; mediodistal facet for the navicular as in A. osborni but subtriangular in outline; ectal facet with concave medial border as in A. osborni and differing from Homalodotherium ; anterodistal end and distal facet little projected in contrast with A. osborni and Homalodotherium ; astragalus with articular head very convex and more medially placed than in A. osborni , separated by a wider neck from the corpus; astragalar groove expanded as in A. osborni ; absence of astragalar foramen as in Homalodotherium ; cuboid relatively wider than in Homalodotherium and without contact with the astragalus; Mc III with proximal facet more extended posteriorly than in Homalodotherium , crescent-shaped and at an acute angle with lateral facets; Mc IV with the groove between medial facets extended to the proximal edge in contrast with Homalodotherium .
Description.—The specimen MCNAM-PV 4958 ( Fig. 7A View Fig ; Table 1) is a large and robust left calcaneum; it is almost complete, only the sustentacular facet is broken at its medial region. The tuber is massive ( Fig. 7A View Fig 1 View Fig ), subtriangular in outline, with a rounded lateral vertex. A well-developed tendinous sulcus crosses from the most posterior border to the anterior apex, separating a lateral and a smaller medial calcaneal processes, the latter more projected proximally ( Fig. 7A View Fig 2 View Fig ). The neck is long and narrow, and its anterior border forms a thin projection between the tuber and the fibular facet, showing a marked semicircular notch at its distal end ( Fig. 7A View Fig 3 View Fig ); this notch could surround a soft structure that, judging by the size, could be a tendon or ligament. The articular area is short and coincides with the holotype of A. petrasnerus (MCNAM-PV 4101; Seoane and Cerdeño 2014); the posterior tuberosity or plantar tubercle (better seen in posterior view; Fig. 7A View Fig 2 View Fig ) is similar to that of the holotype, but is less projected distally than in the specimen MCNAM-PV 3906 (included, but not figured by Seoane and Cerdeño 2014); this difference is interpreted as intraspecific variation. The morphology of MCNAM-PV 4958 differs from both A. osborni (e.g., MACN A 12327, Fig. 7B View Fig ; AMNH 11735) and Homalodotherium (e.g., MACN A 8611, AMNH 9261) in the more developed tendinous sulcus and the projected anterior border with a distal notch, which add to the diagnostic characters of A. petrasnerus . The size of MCNAM-PV 4958 is clearly smaller than that of A. osborni ( Table 1, Fig. 7B View Fig ); this difference in size was not mentioned by Seoane and Cerdeño (2014: fig. 5) and was not reflected in the corresponding figure due to a scale mistake.
Besides MCNAM-PV 4958, a fragment of a left calcaneum, MCNAM-PV 5077, was also recovered. It only preserves the distal part of the sustentacular facet and most of the ectal and fibular facets, which do not differ from the previous specimens of A. petrasnerus ; the slightly greater size of the facets reflects a larger individual.
Stratigraphic and geographic range.—Late Oligocene; Mendoza Province, Argentina.
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