Cecinothofagus Nieves-Aldrey & Liljeblad

Nieves-Aldrey, José Luis, Liljeblad, Johan, Nieves, María Hernández, Grez, Audrey & Nylander, Johan A. A., 2009, Revision and phylogenetics of the genus Paraulax Kieffer (Hymenoptera, Cynipidae) with biological notes and description of a new tribe, a new genus, and five new species, Zootaxa 2200, pp. 1-40 : 18-19

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https://doi.org/ 10.5281/zenodo.189597

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Cecinothofagus Nieves-Aldrey & Liljeblad


Cecinothofagus Nieves-Aldrey & Liljeblad , gen. n.

Type species: Cecinothofagus gallaelenga Nieves-Aldrey & Liljeblad , new species, by present designation

Etymology. An abbreviation alluding to the particle Ceci - (meaning cecidium or gall) and the name of the host plant genus Nothofagus (the southern beeches).

Diagnosis. Differs from Paraulax by the following:

Distinct median vertical carina extending from ventral margin of clypeus to nearly reaching ventral margin of antennal sockets ( Figs. 6 View FIGURE 6 A & 8A). Facial strigae radiating from lateral clypeus, laterally only reaching ventral margin of compound eye. Ventral part of clypeus straight, not projecting over mandibles ( Fig. 6 View FIGURE 6 A). Lateral, sharp occipital carina present. Last antennal flagellomere 1.5 to 1.7 times longer than wide ( Fig. 6 View FIGURE 6 E). F3 of male antenna not modified ( Fig. 6 View FIGURE 6 F).

Longitudinal costulae running from lateral margin of pronotal plate to lateral surface of pronotum: absent or very short ( Fig. 7 View FIGURE 7 A). Notauli sinuate, relatively wider posteriorly, not strongly converging ( Fig. 6 View FIGURE 6 D); distance between notauli at transcutal fissure>0.5 times separation at anterior margin of mesoscutum. Scutellar foveae absent ( Fig. 6 View FIGURE 6 D); scutellum with rugulose-rugose sculpture present marginally but more or less erased in median area. Area above mesopleural impression smooth or weakly coriaceous, usually without longitudinal striae ( Figs. 7 View FIGURE 7 A & 9B). Metascutellum narrower than a metanotal trough in middle. Claws simple, sometimes with a blunt, short basal lobe or with an acute, small lobe measuring less than 1/6 times length of apical tooth ( Fig. 9 View FIGURE 9 D). 3tg of medium size, 0.3–0.5 times length of metasoma. Lateral pubescence on T2 dense. Length of projecting part of hypopygial spine (beyond attachment of lateral flap) <3 times height.

Description. Head. Slightly pubescent; some long, scattered setae on upper frons, vertex and face; some setae also dorsally on occiput, along oral fossa and gena. Gena not expanded behind compound eye. A vertical median carina present ( Figs. 6 View FIGURE 6 A & 8A); facial strigae radiating from clypeus extending laterally, reaching ventral margin of eye; almost absent in median area. Upper face (frons) and vertex usually shining, with delicate, coriaceous sculpture. Clypeus indistinct, ventral margin straight, not projecting over mandibles ( Fig.8 View FIGURE 8 A). Subocular impression present, not well marked. 5–7 regular vertical carinae present ventrolaterally on gena ( Fig. 6 View FIGURE 6 H). Anterior tentorial pits visible; epistomal sulcus and clypeo-pleurostomal lines indistinct. Occiput without dorsal occipital carina; genal carina present ( Fig. 6 View FIGURE 6 B). Hypostomal sulci meeting slightly before hypostoma.

Antenna. Female: 12 segments ( Fig. 6 View FIGURE 6 E); flagellum broadening towards apex; with relatively long, erect setae and placodeal sensilla visible only on flagellar segments F7–F10. Ultimate flagellomere broader than penultimate; more or less spindled-shaped, with a rounded end. Male: 15 segments ( Fig. 6 View FIGURE 6 F). Flagellum not broadening towards apex. F1 cylindrical, F2, and usually also F3, excavated and curved in basal third ( Figs. 6 View FIGURE 6 G & 8D) sometimes apically expanding ( Fig. 4 View FIGURE 4 I). Placodeal sensillae present on all flagellomeres.

Pronotum. Pronotal plate distinct ( Fig. 6 View FIGURE 6 C, 8G), usually without sculpture; dorsal part distinctly set off, anterolateral margin marked and moderately projecting laterad. Admedian pronotal depressions widely separated. Lateral surface of pronotum coriaceous, withouth longitudinal rugae ( Figs. 7 View FIGURE 7 A & 8B).

Mesoscutum almost entirely smooth and shining or with delicate or weak coriaceous sculpture, more marked on lateral lobe ( Fig. 8 View FIGURE 8 A). Some long setae scattered along notauli. Median mesoscutal impression absent ( Fig. 6 View FIGURE 6 D). Notauli percurrent, well separated posteriorly; sometimes ending before transscutal fissure. Anteroadmedian signa visible. Transscutal fissure narrow. Scutellar foveae indistinct, visible only as shallow, usually smooth depression ( Fig. 6 View FIGURE 6 D). Scutellum in dorsal view with weak rugae. Posterodorsal and posterior margins of axillula distinct. Mesopleuron beneath mesopleural triangle with marked, longitudinal mesopleural impression, usually incomplete, not reaching margin of mesopleural triangle ( Fig. 7 View FIGURE 7 A). Area surrounding mesopleural impression almost entirely smooth, withouth visible sculpture. Metascutellum distinctly constricted medially.

Metapectal-propodeal complex. Metapleural sulcus ( Fig. 7 View FIGURE 7 A) meeting posterior margin of mesopectus at about mid height of metapectal-propodeal complex. Lateral propodeal carinae narrow, parallel. Lateral and median propodeal areas smooth, pubescent ( Fig. 7 View FIGURE 7 B). Nucha dorsally with some irregular longitudinal rugae.

Legs. Profemur with ventral swelling in basal third, with 4–5 rows of sharp, closely spaced, deep costulae ( Fig. 7 View FIGURE 7 C). Metatarsal claw simple; sometimes with blunt, short basal lobe or with acute, small lobe less than 1/ 6 of length of apical tooth ( Fig. 9 View FIGURE 9 D).

Forewing. Radial cell closed along anterior margin ( Figs. 7 View FIGURE 7 F & 8H); R1 not or only slightly depigmented along radial cell; areolet absent; vein Rs+M and M weak but visible, directed towards lower half of median vein. Fringe of long setae along apical margin of wing.

Female metasoma laterally compressed ( Fig. 7 View FIGURE 7 D). Abdominal petiole dorsally smooth, ventrally with deep, longitudinal grooves, about as long as high. T2 smooth and shining, covering about 2/3 of metasoma; anteromedian area of T2 with group of long setae. Projecting part of hypopygial spine about 2–3 times longer than high; apical pubescence of hypopigial spine projecting beyond apex, subapical setae longer than apical ones, together forming a small tuft ( Figs. 7 View FIGURE 7 D & 9E).











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