Argia selysi Garrison & von Ellenrieder, 2018

Garrison, Rosser W. & Ellenrieder, Natalia Von, 2018, Damselflies of the genus Argia (Odonata: Coenagrionidae) from Ecuador with descriptions of five new species, Zootaxa 4470 (1), pp. 1-69 : 30-34

Argia selysi Garrison & von Ellenrieder , n. sp.

Figs. 21 (head, thorax, S1– 3 ♂), 32 (head, thorax, S1– 3 ♀), 56 (S7– 10 ♂), 67 (S7– 10 ♀), 83 (mesostigmal plates ♀), 120 (appendages ♂), 140 (map), 149 (photo).

Etymology. Named selysi (Latinized name) after the "Maitre de l'Odonatologie," Baron Edmund de Selys- Longchamps, who, with his friend and collaborator, Herman Hagen, published the first revisionary work on this genus in 1865.

Types. Holotype ♂: ECUADOR, Napo Prov.: Jatun Yacu, Napo River watershed {1°1' S, 77°50' W, 700 m}, 18 April 1935, William Clarke-Macintyre leg. [ UMMZ].

Specimens examined (all paratypes). 418 ♂♂, 136 ♀♀. ECUADOR: Orellana Prov.: 1 ♂, Yasuni National Park, Rosa River, road to Waorani Reserve, 15 km south of main station [coordinates approximate, about 15 airline km south of known coordinates for Station] {0°48'89'' S, 76°22'51'' W, 297 m}, 14 June 1995, Kenneth J. Tennessen leg. [RWG]; 1 ♂, seepages near river, 43.5 km west Loreto, collected under Permit No. 027 1C INEFAN/DNANVS {0°43'0'' S, 77°37'18'' W, 1232 m}, Sidney W. Dunkle leg. [FSCA]; 1 ♂, 1 ♀, small stream near Bigal River Reserve, 30 km north of Loreto (0°33'51'' S, 77°24'41'' W, 880 m), 16 May 2009, Seth Bybee leg. [FSCA]; Pastaza Prov.: 1 ♂, 1.5 km south of Puyo, {1°29'47'' S, 78°0'48'' W, 916 m}, 15–19 May 1977, Paul J. Spangler, Don R. Givens leg. [RWG]; 1 ♂, same data but 19 May 1977 [ RWG]; 1 ♂, 1 ♀, 12 km northeast on road from Puyo to Tena {1°23'45'' S, 77°58'32'' W, 1000 m}, 26 October 1980, Minter J. Westfall, Jr. leg. [ FSCA]; 1 ♂, 16 km northeast on road from Puyo to Tena {1°22'28'' S, 77°57'18'' W, 980 m} [ FSCA]; 5 ♂♂, 24 km northeast on road from Puyo to Tena {1°17'1'' S, 77°53'22'' W, 700 m} [ FSCA]; 2 ♂♂, 33 km northeast on road from Puyo to Tena, 27 October 1980 [ FSCA]; 1 ♂, Abitagua, mouth of Fox River {1°27'0'' S, 78°9'0'' W, 1050 m}, 16 May 1941, William Clarke-Macintyre leg. [ UMMZ]; 1 ♂, Abitagua , Pastaza River watershed {1°27'0'' S, 78°9'0'' W, 1,000 m}, 9 October 1939 [ RWG] ; same data but 1 ♂, 1 ♀, Abitagua, Pastaza River watershed , lower playa, 8 June 1941 [ RWG] ; same data but 2 ♂♂ [ UMMZ]; 4 ♂♂, between Puyo and Mera {1°29' S, 78°3' W}, December 1948 [ RWG] ; same data but 2 ♂♂ [ UMMZ]; 4 ♂♂, between Puyo and Mera [determined as #50 by LK Gloyd and not A. infrequentula ] [ RWG]; 1 ♂, 1 ♀, c-z trail, between Campamento Zatayru and El Partidero {1°22'0'' S, 78°0'0'' W}, 3 August 1935 [ RWG]; 1 ♀, El Partidero-Las Palmas trail {1°22'0'' S, 78°0'0'' W}, 3 August 1935 [ UMMZ]; 1 ♂, 1 ♀, El Partidero, Indian village on east bank of Anzu River , junction of trails from Mera and Puyo northward to Napo {1°22' S, 78°0' W}, 27 July 1935 [ UMMZ] ; same data but 3 ♂♂, 12 November 1936 [ UMMZ]; 1 ♂, El Partidero, Napo River watershed #1 {1°4' S, 77°48' W, 900 m}, 24 August 1935 [ UMMZ]; 1 ♂, 1 ♀, Mera , Pastaza River {1°28' 4" S, 78°6' 14" W, 950 m}, 15 November 1936 [ UMMZ] ; same data but 2 ♂♂ [ UMMZ] ; same data but 1 ♀, 26 February 1946 [ RWG] ; same data but 28 February 1946 [ UMMZ]; 1 ♂, 1 ♀, Pambay {1°28'14'' S, 78°0'21'' W, 945 m}, 15 March 1950 [ UMMZ]; 2 ♂♂, Puyo {1°28' 0'' S, 77°59 '0'' W, 1,000 m}, 5 May 1977, Paul J. Spangler, Don R. Givens leg. [ RWG] ; same data but 1 ♀, 25 April 2009, T. & K. Miyashita leg. [ HK] ; same data but 1 ♂, April 2009 [ HK]; 1 ♂, 1 ♀, Puyo #1, Pastaza River watershed {1°28'0'' S, 77°59'0'' W}, 20 July 1935, William Clarke-Macintyre leg. [ CSCA] ; same data but 2 ♂♂ [ MCZ] ; same data but 1 ♂, 22 July 1935 [ UMMZ] ; same data but 2 ♂♂, 23 July 1935 [ CSCA] ; same data but 2 ♂♂ [ RWG] ; same data but 1 ♀ [ UMMZ] ; same data but 1 ♂ [ UMMZ] ; same data but 1 ♂, 24 July 1935 [ UMMZ] ; same data but 1 ♂, 30 July 1935 [ UMMZ] ; same data but 2 ♂♂ [ UMMZ] ; same data but 1 ♂, July 1935 [ UMMZ]; 2 ♂♂, Puyo #2, Pastaza River watershed {1°28'0'' S, 77°59'0'' W}, 21 July 1935 [ BMNH] ; same data but 1 ♀ [ CSCA] ; same data but 2 ♂♂ [ CSCA] ; same data but 1 ♀ [ RWG] ; same data but 2 ♂♂ [ RWG] ; same data but 1 ♂, 1 ♀ [ UMMZ] ; same data but 2 ♀♀ [ UMMZ] ; same data but 8 ♂♂ [ UMMZ] ; same data but 1 ♂, 22 July 1935 [ UMMZ] ; same data but 4 ♂♂, 26 July 1935 [ UMMZ] ; same data but 1 ♂ [ UMMZ] ; same data but 2 ♂♂, 27 July 1935 [ CSCA] ; same data but 2 ♂♂ [ RWG] ; same data but 2 ♂♂ [ UMMZ]; 1 ♂, Puyo #3, Pastaza River watershed {1°28'0'' S, 77°59'0'' W}, 20 August 1935 [ CSCA] ; same data but 1 ♀ [ CSCA] ; same data but 1 ♂ [ RWG] ; same data but 1 ♀ [ RWG] ; same data but 3 ♂♂ [ UMMZ] ; same data but 4 ♀♀ [ UMMZ]; 1 ♂, Puyo to Pastaza , Pastaza River watershed {1°28'0'' S, 77°59'0'' W, 981 m}, December 1948 [ RWG] ; same data but 1 ♂, 6 March 1950 [ RWG] ; same data but 1 ♂, March 1950 [ BMNH] ; same data but 1 ♂, December 1948 [ MCZ] ; same data but 1 ♂ [ RWG] ; same data but 2 ♂♂, 1 ♀, 1949 [ UMMZ]; 1 ♂, Napo River near Puyo , Pastaza River watershed {1°28'0'' S, 77°59'0'' W}, 20 November 1936 [ UMMZ] ; same data but 1 ♂, 1 ♀, 28 November 1936 [ UMMZ] ; same data but 1 ♂, 25 November 1935 [ CSCA] ; same data but 1 ♀ [ CSCA] ; same data but 1 ♂ [ RWG] ; same data but 4 ♂♂ [ UMMZ] ; same data but 1 ♂, 29 November 1935 [ UMMZ]; 1 ♀, Yanamanaca [next Indian village up Anzu River from Yanamanaca is El Partidero; coordinates for El Partidero], Napo River watershed {1°22' S, 78°0' W}, 11 September 1935 [ RWG] ; same data but 1 ♂ [ RWG] ; same data but 2 ♂♂ [ UMMZ] ; same data but 1 ♀, 1 ♂ 20 September 1935 [ CSCA] ; same data but 2 ♂♂, 1 ♀ [ RWG] ; same data but 7 ♂♂, 3 ♀♀ [ UMMZ] ; same data but 2 ♂♂, 1 ♀, 29 September 1935 [ CSCA] ; same data but 2 ♂♂, 1 ♀ [ RWG] ; same data but 15 ♂♂, 6 ♀♀ [ UMMZ] ; same data but 10 ♂♂, 3 ♀♀, 17 October 1935 [ UMMZ] ; same data but 6 ♂♂, 2 ♀♀, 2 November 1935 [ CSCA] ; same data but 6 ♂♂, 2 ♀♀ [ RWG] ; same data but 19 ♂♂, 6 ♀♀ [ UMMZ] ; same data but 3 ♂♂, 3 ♀♀ [ UMMZ]; 2 ♀♀, Zulay, Pastaza River watershed {1°29'26'' S, 78°3'43'' W}, 14 March 1950 [ RWG] ; same data but 3 ♀♀ [ UMMZ]; Napo Prov.: 1 ♂, 1 ♀ (in copula), 8.3 km west of Pano [determined by Gloyd as sp. #50] {1°1'35'' S, 77°53'11'' W, 590 m}, 22 October 1976, Herman G. Real leg. [ RWG]; 1 ♀ , same data but along trail between main road (Puyo-Baños) and Bombaini- Yacu, Pastaza River watershed {1°30' S, 78°0' W, 400 m}, 21 February 1950, William Clarke-Macintyre leg. [ RWG]; 1 ♂, Boya Yacu, Napo watershed River {1°1'0'' S, 77°50'0'' W, 700 m}, 3 September 1935 [ UMMZ]; 1 ♂, Chonta-yacu {1°1'0'' S, 77°50'0'' W, 900 m} [ UMMZ]; 1 ♂, Concepción, Napo River {1°40' S, 77°25' W, 400 m}, 28 December 1939 [ UMMZ]; 1 ♂, east branch of Ishpinga-yacu, Napo {0°59'20'' S, 77°50'12'' W, 500 m}, 18 September 1942 [ UMMZ]; 1 ♂, Gavilanes, Napo River watershed {1°1'0'' S, 77°50'0'' W, 700 m}, 20 September 1935 [ UMMZ]; 1 ♂, Hacienda Ila , Anzu River {1°12' S, 77°50' W}, 6 August 1935 [ CSCA] ; same data but 10 ♂♂, 3 ♀♀, 16 December 1936 [ UMMZ] ; same data but 1 ♂, 2 ♀♀, 17 December 1936 [ CSCA] ; same data but 4 ♂♂, 2 ♀♀ [ UMMZ] ; same data but 2 ♂♂, 1 ♀, 18 December 1936 [ UMMZ]; 2 ♂♂, Headwaters of Churi-yacu, Pastaza River watershed {1°2'0'' S, 77°41'0'' W}, 900 m, 29 May 1941 [ UMMZ]; 2 ♂♂, Headwaters of Arajuno River, Napo watershed, stream #1 {1°4'46'' S, 77°32'28'' W, 1,000 m}, 22 April 1941 [ CSCA] ; same data but 3 ♂♂, 1 ♀ [ RWG] ; same data but 6 ♂♂, 1 ♀ [ UMMZ] ; same data but 5 ♂♂, 1 ♀, 23 April 1941 [ UMMZ] ; same data but 1 ♂, 24 April 1941 [ UMMZ] ; same data but 3 ♂♂, 26 April 1941 [ UMMZ] ; same data but 2 ♂♂, 1 ♀, 27 April 1941 [ CSCA] ; same data but 2 ♂♂ [ UMMZ] ; same data but 1 ♀, 29 April 1941 [ UMMZ] ; same data but 2 ♂♂, 1 ♀, 30 April 1941 [ RWG] ; same data but 1 ♂, 1 ♀ [ UMMZ]; 2 ♂♂, Ishpinga-yacu {0°59'20'' S, 77°50'12'' W, 500 m}, 12 August 1942 [ UMMZ]; 1 ♂, 1 ♀, Ishpinga-yacu, Upper Napo River {0°59'20'' S, 77°50'12'' W, 500 m}, 1 September 1942 [ RWG]; 1 ♂, Jatun Yacu , Napo River watershed {1°1'0'' S, 77°50'0'' W, 700 m}, 5 April 1935 [ UMMZ] ; same data but 1 ♂, 11 April 1935 [ UMMZ] ; same data but 1 ♂, 17 April 1935 [ UMMZ] ; same data but 1 ♀, 18 April 1935 [ CSCA] ; same data but 1 ♂ [ RWG] ; same data but 3 ♂♂ [ UMMZ] ; same data but 1 ♀, 7 February 1937 [ RWG] ; same data but 6 ♂♂ [ UMMZ] ; same data but 1 ♂, 1 ♀, 12 February 1937 [ RWG] ; same data but 2 ♂♂, 1 ♀, 24 February 1937 [ RWG] ; same data but 1 ♂ [ UMMZ] ; same data but 1 ♂ 25 February 1937 [ RWG] ; same data but 2 ♂♂, March 1937 [ BMNH] ; same data but 6 ♂♂, 6 ♀♀ [ CSCA] ; same data but 2 ♂♂ [ MCZ] ; same data but 1 ♂, March 1940 [ RWG] ; same data but 2 ♀♀, 3 July 1937 [ RWG] ; same data but 1 ♂, 1 ♀, 4 July 1937 [ RWG] ; same data but 6 ♂♂, 6 ♀♀, 3 July 1937 [ RWG] ; same data but 4 ♂♂, 4 July 1937 [ UMMZ] ; same data but 22 ♂♂, 7 ♀♀, 3 July 1937 [ UMMZ]; 1 ♀, Jatun Yacu, vicinity of Napo {1°1'0'' S, 77°50'0'' W}, 1 November 1934 [ RWG]; 1 ♀, Las Palmas , on Anzu River ( Napo River watershed ) {1°4' S, 77°48' W, 900 m}, 16 August 1935 [ RWG] ; same data but 1 ♂ [ UMMZ] ; same data but 1 ♂, 17 August 1935 [ RWG] ; same data but 1 ♀ [ UMMZ]; 1 ♂, Las Palmas, on Anzu River ( Napo River watershed ), in yard{1°4' S, 77°48' W}, 16 August 1935 [ UMMZ]; 6 ♂♂, Las Palmas , Anzu River , Napo River watershed {1°4' S, 77°48' W}, 11 December 1936 [ CSCA] ; same data but 6 ♂♂ [ RWG] ; same data but 5 ♂♂, 5 ♀♀ [ UMMZ] ; same data but 1 ♂ [ UMMZ]; 1 ♀, Las Palmas, Anzu River , Napo River watershed #1 {1°4' S, 77°48' W} 16 August 1935 [ UMMZ] ; same data but 2 ♂♂, 2 ♀♀, 11 December 1936 [ UMMZ]; 1 ♂, Llandia Hill, Napo River watershed {1°1'0'' S, 77°50'0'' W, 700 m}, 15 September 1935 [ UMMZ]; 4 ♂♂, north side Bombaini-Yacu River , Pastaza River watershed {1°30' S, 78°0' W, 900 m}, 24 March 1941 [ RWG]; 1 ♂, Napo-Puyo road (1°23'29'' S, 77°51'35'' W, 609 m), 6 November 2001, Fred C. Sibley leg. [ FSCA]; 5 ♂♂, 1 ♀, pond and small stream 34 km west Loreto, collected under Permit No. 027 1C INEFAN/DNANVS {0°43'30'' S, 77°34'31'' W, 840 m}, 14 June 1995, Sidney W. Dunkle leg. [ RWG]; 1 ♂, Llandia River , Napo River watershed {1°1'0'' S, 77°50'0'' W, 700 m}, 2 September 1935, William Clarke-Macintyre leg. [ UMMZ]; Anzu River (vicinity of Napo ) {1°3'9'' S, 77°48'12'' W, 900 m}, 7 July 1934 [ RWG] ; same data but 1 ♂, 1 ♀, 19 July 1934 [ RWG] ; same data but 1 ♂, 26 July 1934 [ RWG] ; same data but 1 ♀, 4 August 1934 [ RWG] ; same data but 1 ♂, 1 ♀, 6 August 1934 [ RWG] ; same data but 1 ♀, 9 August 1934 [ RWG] ; same data but 1 ♀, 12 August 1934 [ RWG]; 2 ♂♂, Bombaini-Yacu River , Pastaza River watershed {1°30' S, 78°0' W}, 19 March 1941 [ CSCA] ; same data but 2 ♂♂ [ RWG] ; same data but 14 ♂♂, 3 ♀♀ [ UMMZ] ; same data but 1 ♂, 22 March 1941 [ UMMZ] ; same data but 1 ♂, 24 March 1941 [ UMMZ] ; same data but 2 ♂♂, 1 ♀, 12 May 1941 [ CSCA] ; same data but 2 ♂♂, 1 ♀ [ RWG] ; same data but 2 ♂♂, 1 ♀ [ UMMZ]; 1 ♂, Ila River (near Hacienda Ila ) {1°12' S, 77°50' W, 700 m}, 14 August 1934 [ RWG] ; same data but 1 ♀, 18 August 1934 [ RWG] ; same data but 1 ♂, 1 ♀, 3 November 1934 [ RWG]; 3 ♂♂, seep near large rocky river, 43.5 km west of Loreto (0°43'16'' S, 77°38'28'' W, 880 m), 14 June 1995, William F. Mauffray leg. [ FSCA]; 1 ♂, seeps/small stream near river, 5 km west of Orellana Province border (12 km west of Huiruno ), highway 45; 2009-0186 (0°44'22'' S, 77°33'11'' W, 940 m), 11 October 2009, Jim T. Johnson leg. [ JJ]; 1 ♂, Upper Anzu River , Napo {1°4' S, 77°48' W, 500 m}, August 1942, William Clarke-Macintyre leg. [ UMMZ]; 1 ♀, Upper Ila River {1°12' S, 77°50' W, 700 m}, 21 November 1934 [ RWG]; 1 ♂, Yurayacu , Napo River watershed {1°1'0'' S, 77°50'0'' W}, 17 September 1935 [ UMMZ] ; same data but 1 ♂, 1 ♀, 27 September 1935 [ UMMZ]; 1 ♂, 1 ♀, Zatzayacu, Anzu River {1°3'9'' S, 77°48'12'' W, 1000 m}, 17 December 1936 [ RWG] ; same data but 3 ♂♂, 1 ♀ [ UMMZ]; 2 ♂♂, Zatzayacu, Anzu River, Napo River watershed {1°3'9'' S, 77°48'12'' W}, 24 August 1935 [ UMMZ]; Tungurahua Prov. : 1 ♂, stream, highway 50 between Verde River and Negro River, 2009-0109 (1°24'25'' S, 78°15'19'' W, 1,515 m), 4 October 2009, Jim T. Johnson leg. [ JJ]; Morona Santiago Prov. : 8 ♂♂, 57 km north on highway 45 (80 km north of Sucua ), Pasanac River , seepages {1°56'53'' S, 77°51'11'' W, 780 m}, 10 November 1997, William F. Mauffray leg. [ FSCA] ; same data but 2 ♂♂ [ RWG]; 7 ♂♂, 58 km north on highway 45 (70 km north of Sucua), roadside ditch and tributaries near Pasanac River , 16 September 2005 [ FSCA] ; same data but 1 ♂ [ RWG]; 3 ♂♂, Macas, 1 km south of Pasanac River (56 km north on highway 45), small stream in open grassy field (1°57'0'' S, 77°51'46'' W, 213 m) [ FSCA]; 10 ♂♂, Macas, 57 km north on highway 45 (80 km north of Sucua ), Pasanac River , seepages (1°56'41'' S, 77°51'13'' W, 782 m), 10 November 1997 [ FSCA]; 1 ♂, Mangosisa , Upano River , Received November 30, 1945 {2°31'28'' S, 77°53'41'' W, 850 m}, Leopoldo Gomez Alonzo leg. [ RWG]; 1 ♂, roadside ditch 22.5 km north of Indanza , highway 45 (2°55'9'' S, 78°24'5'' W, 1,110 m), 7 November 1997, William F. Mauffray leg. [ FSCA]; 1 ♂, 1 ♀ (tandem pair), small stream about 1 km south of Pasanac River , highway 45;(1°57'0'' S, 77°51'46'' W, 700 m), 16 September 2005, Kenneth J. Tennessen leg. [ FSCA]; 1 ♂, small swamp 7 km south of Comunidad Chuwitayo {1°57'2'' S, 77°51'4'' W}, 9 November 1997, Thomas W. Donnelly leg. [ RWG]; 2 ♂♂, waterfall/seepage pond by Pasanac River , 80 km north of Sucua , noon {1°57'13'' S, 77°51'27'' W, 650 m}, 10 November 1997, William F. Mauffray leg. [ RWG] .

A moderate-sized dark species of the Argia huanacina -group with reduced pale colors dark purple, welldeveloped pale occipital bar, and small, rounded cercus with a small apicomedial tooth and trilobate paraproct.

Description of male holotype. Head: Labium, ante- and postclypeus, genae and ante- and postfrons entirely pale (these pale colors darkened due to postmortem affects), epicranium black with small pale spot anterolateral to lateral ocellus, black appearing as a broad cross-band delimited posteriorly by large purple postocular spots continuous with pale occipital bar (as in Fig. 21 ), dorsal surface of postocular lobe black; antennae black, rear of head black with lateral border along eye pale (as in Fig. 5 ).

Prothorax black with following areas violaceous: large mediolateral spot on middle lobe, lateral 0.20 of hind lobe, and propleuron (as in Fig. 21 ). Pale areas of synthorax violet, with broad black middorsal stripe about twice as wide as each pale parallel antehumeral stripe; parallel black humeral stripe extending from base of mesinfraepisternum and enclosing a small pale spot at dorsal 0.10, a narrow dark brown stripe over interpleural suture separating pale areas on side of thorax (as in Fig. 21 ). Wings hyaline, with venation black; pterostigma brown, surmounting 1 cell in all wings; postnodals Fw 14/15, Hw 12/12; postquadrangular cells Fw 3/3, Hw 3/3; RP 2 at Fw 7/8, Hw 6/6. Coxae pale; trochanters pale anteriorly, brown posteriorly; femora black externally, pale internally; tibiae pale externally black laterally and internally; tarsi and armature black.

Abdomen (as in Figs. 21 , 56) violaceous and black; S1 pale with small black spot dorsally on basal 0.50; S2 dorsally with pale spot narrowing posteriorly ending before apical margin, ventral margin of S2 pale, with a lateral longitudinal black stripe broadening posteriorly and meeting dorsally at posterior 0.20 of segment and connected apically by black annular ring; S3 with basal ring of violet, remainder black except for obscure pale green ventrolateral stripe at along medial half of segment; S4–7 as in S3 but with pale areas successively reduced; S8–10 violet except for narrow edge of black on posterolateral margin of S10; cercus brown, paraproct black.

Genital ligula as illustrated for Argia pulla ( Fig. 9u in Garrison & von Ellenrieder 2015), with distal segment ending in two filaments which at their base are covered by a narrow, long digit-like membranous hood and with a recurved sclerotized lateral lobe just distal to flexure of genital ligula on each side.

Torus large, transversely triangular, swollen ventrally, occupying entire ventral margin of torifer, almost approximate and partially overlapping bilobed epiproct ( Fig. 120b ), base of epiproct and surrounding area black; cercus in dorsal view roundly quadrate, slightly longer medially than laterally and armed with a small recurved tooth at apicomedial margin ( Figs. 120c, d ); paraproct in lateral view trilobate with lobes acuminate ( Fig. 120a ), the upright dorsal lobe the largest with posterior margin at a right angle to second distal lobe in lateral view.

Dimensions of holotype. Hw 18.0, abdomen 26.0, total length 32.0.

Description of female paratype ( Ecuador: Napo Prov., Zatzayacu, Anzu River). Head, pro-, synthorax and S1–2 as in male ( Fig. 32 ) but pale prothoracic middorsal spots on middle lobe conjoined dorsally; S3–6 as in male but pale areas slightly more extensive, pale ventrolateral areas on S3–6 larger and of definite outline, S7 as in male but with pale ventral stripe wider, S8 violet dorsally with irregular black lateral stripe narrowing at distal 0.30 before widening, anterior 0.50 of black stripe separated ventrally by narrow pale stripe; S9 pale with anteroposteriorly directed elongate black spot at basal 0.50, its end not connected above and with disjunct black ventral stripe widening posteriorly; S10 black with irregular pale lateral spot, ovipositor pale (Fig. 67).

Mesostigmal lobe as in A. huanacina ( Fig. 76 ), small, digit-like, raised medially and with outer margin of lobe curved ventrally giving appearance of a notch at tip of lobe ( Figs. 83a, b, d ); a shallow semicircular emarginate depression on mesepisternum immediately posterior to mesostigmal lobe ( Fig. 83d ).

Variation in paratypes. Little variation was observed in the series collected at the type locality. Pterostigma surmounting 1 cell in males and females; postnodals: Fw 14–15, Hw 12–14 in males, Fw 13–15, Hw 12–13 in females; postquadrangular cells Fw 3, Hw 3 in males, Fw 3–4 (usually 3), Hw 3 in females; RP 2 at Fw 8, Hw 6–7 in males, Fw 7–8, Hw 6–7 in females.

Dimensions of paratypes. ♂: Hw 17.8 ± 0.55 [16.7–18.5], abdomen 25.1 ± 0.98 [23.4–26.5], total length 31.4 ± 1.14 [29.4–32.7]. ♀: Hw 18.3 ± 0.44 [17.7–18.9], abdomen 24.8 ± 0.66 [23.6–25.7], total length 31.1 ± 1.01 [29.6–32.9].

Diagnosis. This is the fourth species belonging to the Argia huanacina group (and mentioned as such in Garrison & von Ellenrieder 2015: 25) characterized in the male by the acutely trilobate cercus in lateral view and small curved digitiform mesostigmal lobe in the female. The male is most similar to Argia huanacina and will key to that species in Garrison & von Ellenrieder (2015) differing by having the mesal margin of cercus linear and longer than outer margin in dorsal view ( Fig 120b ; mesal margin of cercus convex and subequal to outer margin in A. huanacina , Fig. 110b ) and by possessing a recurved tooth at mesodistal margin of cercus ventrally, Fig. 120d (similar tooth along mesal margin in A. huanacina , Fig. 110d ).

Female is most similar by mesostigmal lobe morphology to Argia huanacina and their separation requires care. Anterior part of mesepisternum directly behind mesostigmal lobe is depressed and shallowly emarginate in A. selysi , Fig. 83a (no differentiation of mesepisternum directly behind mesostigmal lobe in A. huanacina , Fig. 76a ). The largely allopatric distribution ( A. selysi restricted to Ecuador, A. huanacina with one record from Ecuador, otherwise in Peru, Bolivia, and Brazil, Fig. 140 ) will further aid in separating these two closely related species. Argia selysi is broadly sympatric with A. infrequentula Fraser ( Fig. 140 ).

Both sexes of Argia selysi key to Argia huanacina in Garrison & von Ellenrieder (2015) from which the last couplet (2') may be modified as follows:

Figures

FIGURES 19–22. Color pattern of male head, thorax, and S1–4. Head, dorsal view, thorax and S1–4, lateral view.

FIGURES 1–8. Color pattern of head. Head, anterior view (1); head posterior view (2–5); labrum, anterior view (6–8).

FIGURES 9–13. Color pattern of male head, thorax, and S1–4. Head, dorsal view, thorax and S1–4, lateral view.

FIGURES 119–122. Male caudal appendages. (a) lateral view; (b) dorsal view; (c) mediodorsal view; (d) ventral view.

FIGURES 28–33. Color pattern of female. Head, thorax, and S1–4 (31–33); pterothorax (28–30). Head, dorsal view, thorax and S1–4, lateral view.

FIGURES 73–76. Female mesostigmal plates. (a) dorsal view; (b) detail of mesostigmal lobe, dorsal view; (c) frontal (anterior) view; (d) detail of mesostigmal lobe, anteromedial view; (e) laterodorsal view; (f) detail of mesostigmal lobe, anterior view; (g) detail of mesostigmal lobe, posterior view; (h) posterior view.

FIGURES 77–83. Female mesostigmal plates. (a) dorsal view; (b) laterodorsal view; (c) lateral view; (d) detail of mesostigmal lobe, dorsal view; (e) detail of mesostigmal lobe, medial view; (f) frontal (anterior) view; (g) detail of mesostigmal lobe, posterior view; (h) posterior view.

FIGURES 107–110. Male caudal appendages. (a) lateral view; (b) dorsal view; (c) mediodorsal view; (d) posterior view.

FIGURES 140–141. Distribution of new Argia species and closest congeners in Ecuador.