Theroscopus maruyamanus ( Uchida, 1930 )

Watanabe, Kyohei, 2022, Distribution Records and Re-descriptions of Some Japanese Species of the Subfamily Phygadeuontinae (Hymenoptera, Ichneumonidae), Bulletin of the Kanagawa Prefectural Museum (Natural Science) 51, pp. 61-72 : 70-71

Theroscopus maruyamanus ( Uchida, 1930) (SJN: Kitaguni-chibi-togari-himebachi)

( Figs 6A–E )

Phygadeuon maruyamanus Uchida, 1930: 336 .

Description. Female (n=1). Body length 5.9 mm. Body covered with silver setae.

Head matt, 0.6 times as long as wide in dorsal view. Clypeus 0.45 times as wide as maximum length, sparsely punctate dorsally, its anterior margin with a pair of minute median teeth. Frons with smooth areas above antennal sockets. Length of malar space 1.1 times as long as basal width of mandible. OD: POL: OOL = 0.5: 0.8: 0.95. Occipital carina complete, its lower end joined with hypostomal carina distant from mandibular base. Upper tooth of mandible slightly longer than lower tooth. Base of mandiblE flat, with a shallow concavity. AntEnna brokEn in KPM-NK 81838, see remarks.

Mesosoma polished. Lateral part of pronotum punctate dorsally, longitudinally rugulose along posterior margin. Epomia present. Mesoscutum punctate, matt medially. Notaulus weak and short. Scutellum densely punctate, without a lateral longitudinal carina except for base. Mesopleuron covered with fine longitudinal striae and fine punctures except for smooth area on speculum and its anterior area. Epicnemial carina present laterally, its dorsal end not reaching anterior margin of mesopleuron. Sternaulus present except for posterior 0.45. Posterior transverse carina of mesosternum largely absent in front of middle coxa. Metapleuron punctate, with a complete juxtacoxal carina. Propodeal carinae and areas complete. Postero-lateral corner of area densipara projected. Propodeum punctate except for area superomedia largely smooth. Area basalis slightly narrower than area superomedia. Area superomedia as long as wide, receiving lateral section of anterior transverse carina just behind of middle ( Fig. 6E ). Length of fore wing 4.9–6.0 mm. Areolet absent. Vein 1cu-a interstitial to vein M&RS. Nervellus subvertical, intercepted posterior 0.75. Hind femur 4.0 times as long as maximum depth in lateral view. Hind TS I: II: III: IV: V = 2.0: 1.0: 0.7: 0.3: 0.7. Tarsal claws simple.

Metasoma polished. T I 1.5 times as long as maximum width, coriaceous anteriorly, longitudinally striate posteriorly. Median dorsal carina of T I present except for posterior part. Dorsolateral carina of T I complete except it on spiracle. T II 0.55 times as long as maximum width, coriaceous except for smooth posterior area. T III to T VII polishEd and finEly punctatE. Ovipositor shEath 0.8 timEs as long as hind tibia. Ovipositor straight, with a nodus and ventral teeth.

Coloration ( Figs. 6A–C, E ). Body (excluding wings and lEgs) black to blackish brown (pEdicEl and flagEllum see remarks below). Basal spot of mandible yellowish brown. Tegula brown. T II, T III, and posterior margins of T IV and T V reddish brown. Posterior parts of T VI and T VII white. Membranous part of metasomal sternites and ovipositor reddish brown. Wings hyaline. Veins and pterostigma brown to blackish brown except for base of pterostigma yellowish brown. Legs reddish brown to dark reddish brown. Coxae and trochanters black. Hind trochantellus, femur, base and apical part of tibia, and tarsus darkened.

Male. Unknown.

Material examined. JAPAN: KPM-NK 81838, F, Tokyo, Hinohara Vil., Mt. Outakyama , 1. VI . 2008, T. Ban leg.

Distribution. Japan (Hokkaido and Honshu).

Bionomics. Unknown.

Remarks. KPM-NK 81838 has been compared with the lectotype deposited in SEHU. Although the antennae of KPM-NK 81838 are lost, I conclude that this specimen is T. maruyamanus by the other characteristics. KPM-NK 81838 is thE sEcond rEcord of this spEciEs and is thE first record of this species from Honshu. The character states of thE antEnna of lEctotypE: antEnna with 20 flagEllomErEs, subapical part slightly widened; length of FL I 2.7 times as maximum depth of FL I in lateral view and 1.0 times as long as length of FL II; FL IV to FL VII white ( Fig. 6D ). KPM-NK 81838 differs from the lectotype in the following three characteristics: body length 5.9 mm (7.0 mm in lectotype); T I 1.5 times as long as maximum width (1.3 times in lectotype); ovipositor sheath 0.8 times as long as hind tibia (ca. 0.55 times in lectotype), while I judged that thEsE charactEr statEs arE intraspEcific variations.

Figures

Fig. 6.Theroscopus maruyamanus (Uchida, 1930), females (A–C, E: KPM-NK 81838; D: lectotype deposited in SEHU) ― A: dorsal habitus; B: lateral habitus; C: head, frontal view; D: head and antenna, lateral view; E: scutellum and propodeum, dorsal view.