Heynea multijuga Blume, Bijdr. Fl. Ned. Ind.

Holzmeyer, Laura, Hauenschild, Frank, Mabberley, David J. & Muellner-Riehl, Alexandra N., 2021, Confirmed polyphyly, generic recircumscription and typification of Dysoxylum (Meliaceae), with revised disposition of currently accepted species, TAXON 70 (6), pp. 1248-1272 : 1258

Heynea multijuga Blume, Bijdr. Fl. Ned. Ind. : 168. 1825

= Dysoxylum cyrtobotryum Miq., Fl. Ned. Ind., Eerste Bijv. : 196, 504. 1861.

Dysoxylum in its new form is essentially Dysoxylum sect. Cyrtochiton , as circumscribed by Mabberley (1995b), and is represented in the phylogenetic tree by clade 6 ( Fig. 1B ). Didymocheton , Epicharis , Goniocheton , Prasoxylon , and Pseudocarapa are represented by clade 1 ( Fig. 1B ), clades 4 & 5 ( Fig. 1B ), clade 8 ( Fig. 1B ), clade 11 ( Fig. 1A ) and clade 7 ( Fig. 1B ), respectively.

This generic resolution is a necessary precursor to detailed monographic work on the genera here recognized. Names are very urgently needed (cf. Heads, 2019) in the countries where these important trees are native and often some of the most frequently encountered and collected. Therefore, a conspectus based on published floristic accounts, which have full descriptions, synonymy and typifications, is presented here. Of the 94 accepted species, those investigated in each recognized genus in this molecular study are listed first, followed by those species that were not. In the interests of conciseness and avoidance of repetition, the descriptions and isotypes and most synonyms of the accepted species are excluded, unless they have not been fully addressed in the recent floristic literature also cited here, so as to allow ready access to the germane literature until definitive monographs are written. Names long-excluded from Dysoxylum s.l. are omitted altogether, as they are also accounted for in that literature.

Figures

Fig. 1. Phylogenetic tree based on BI analysis of the four-marker dataset (ITS, ETS, trnL-F, rps15-ycf1) with two runs of six MCMC chains of 50 million generations and corresponding posterior probability values. In square brackets, we provide the previous species names for all newly resurrected genera. Accession numbers in the tree correspond to specimens listed in Appendix 1. A, Clade 10 (green) represents all accessions of Chisocheton referred to as “Ch.”; clade 11 (pink) shows all accessions of Prasoxylon referred to as “Pra.”; clade 12 (orange) shows all accessions of Anthocarapa referred to as “An.”, and Synoum as “Sy.”; outgroup (white/no colour) with Trichilia as “Tr.”, Cipadessa as “Ci.”, Nymania as “Ny.” and Turraea as “Tu.”. B, Clade 1 (yellow) represents all accessions of Didymocheton referred to as “D.”; clade 2 (dark blue) shows Cabralea referred to as “C.”; clade 3 (green) represents Aglaieae sensu Pennington & Styles, containing Aglaia referred to as “Ag.”, Aphanamixis as “A.”, Reinwardtiodendron as “Re.”, Lansium as “La.”, and Sphaerosacme as “Sph.”; clades 4 and 5 (pink) show all accessions of Epicharis referred to as “E.”; clades 6 (light blue), 7 (purple) and 8 (red) show all accessions of Dysoxylum s.str. referred to as “Dys.”, Pseudocarapa referred to as “Ps.”, and Goniocheton referred to as “G.”, respectively; clade 9 (orange) with the large parts of ‘Guareeae’ represents all accessions of Heckeldora referred to as “H.”, Guarea as “Gu.”, Leplaea as “Le.”, Neoguarea as “Neog.”, Ruagea as “Ru.”, and Turraeanthus as “Tur.”.