Titanogrypa, , Pape, 1996

Barbosa, Taciano M., Mello-Patiu, Cátia A. & Vasconcelos, Simão D., 2021, Revision of the New World subgenus Titanogrypa (Cucullomyia) (Diptera: Sarcophagidae), with a reassessment of diagnostic characters, Journal of Natural History 55 (5 - 6), pp. 305-340 : 309-310

publication ID	https://doi.org/ 10.1080/00222933.2021.1902587
persistent identifier	https://treatment.plazi.org/id/03AF87CE-3E7A-2B40-F1A9-FD2FB38FF905
treatment provided by	Plazi
scientific name	Titanogrypa
status

Treatment

Titanogrypa View in CoL View at ENA ( Cucullomyia Roback, 1954 )

Cucullomyia Roback 1954: 83 . Type species: Sarcophaga pedunculata Hall, 1931 View in CoL (by original designation). Titanogrypa View in CoL ; Downes 1965: 956.(new synonym). Cuculomyia ; Lopes 1969: 47. [incorrect subsequent spelling of Cucullomyia ] (catalogue). Cucullomyia ; Lopes 1976b (revision, key and new species descriptions); Lopes 1982: 316. (larva 1 description, subtribe); Lopes 1988: 101. (new species description). Titanogrypa (Cucullomyia) View in CoL ; Pape 1996: 431. (catalogue).

Generic diagnosis. Frontal row of setae not diverging near lunule, male without proclinate orbital setae; gena with dark anterior and pale posterior setae; four post-sutural dorsocentral setae, the two posterior ones larger; postalar wall setulose; scutellum with a patch of hair-like setulae on lateral margins, reaching ventral part, black in T. (C.) luculenta and T. (C.) ecuatoriana , whitish in remaining species; three pairs of marginal scutellar setae (basal, lateral and subapical), one pair of discal scutellar and no apical scutellar setae (except in T. (C.) luculenta ); third costal sector without ventral setulae; mid femur with apical posteroventral ctenidium (absent in T. (C.) luculenta ); ventral margin of T3–T5 with long hair-like setae with curling apex, more numerous on T5; T4 with a pair of median marginal setae; ST5 of the male deeply and widely cleft as an inverted V, with a short base without a window and narrow arms with hair-like and spine-like setae; male terminalia with syntergosternite 7 + 8 and epandrium reddish brown in ground colour; pregonite long and spatulated; cercus with broader proximal half and apex with characteristic shape, distal end slightly enlarged and rounded in profile, except in T. albuquerquei which presenting the cercus distal end slightly tapered and curved in profile; cercal prongs parallel in posterior view; phallus with a distinct hinge between basi- and distiphallus; basiphallus with a dorsal hump at junction with distiphallus; basiphallus long (at least 2.5x phallic tube length), slender, and laterally compressed, with a longitudinal keel on dorsal face; paraphallus short with a dorsal pair of keels; vesica conspicuously developed forwards and forming a hood over the styli; median and lateral styli long and tubular, with distinct openings; juxta distinct, like a pair of separate lobes.

Key to males of Titanogrypa (Cucullomyia) View in CoL

1. Scutellum with a lateral patch of whitish or yellowish hair-like setulae ( Figure 7 View Figure 7 (c)); pregonite with base and apex similar in width; and surstylus clavate, sometimes with short projection on posterior margin........................................................................................................... 2

1 ʹ. Scutellum with a lateral patch of black hair-like setulae ( Figure 7 View Figure 7 (d)); pregonite with broad base and narrow apex; and surstylus triangular and boomerang-like, with anterior margin projected forward ( Figure 5 View Figure 5 (b))................................................................................................ 6

2. Vesica without any upward projections at its base ( Figure 6 View Figure 6 (e)), elongate, with a sclerotised sheath along distal margin; juxtal lobes broad, as long as paraphallus ( Figure 6 View Figure 6 (c,e)).................................................... .................................................... T. (C.) placida (Aldrich) View in CoL

2 ʹ Vesica with a membranous or sclerotised upward projection, like a tip or flap ( Figure 1 View Figure 1 (e)); juxtal lobes shorter than paraphallus........................................................................................................... 3

3. Vesica as long as wide, with a single sclerotised proximal projection at base and a rounded and spinous distal portion ( Figure 4 View Figure 4 (d,e))............. ............. T. (C.) larvicida (Lopes)

3 ʹ. Vesica 2x as longer as wide, with one or more membranous projections and without ornamentation with spines in distal portion ( Figure 2 View Figure 2 (e))............................................................ 4

4. Cercal prong with distal end slightly tapered and curved in profile ( Figure 2 View Figure 2 (b)); vesica with a proximal membranous projection and two distal projections as lateral flaps ( Figure 2 View Figure 2 (d,e))... ............................................................................................................................................ T. (C.) albuquerquei (Lopes) View in CoL

4 ʹ. Cercal prong with distal end slightly enlarged and rounded in profile ( Figure 1 View Figure 1 (b)); vesica with only the proximal membranous projection, without distal projections ( Figures 1 View Figure 1 (e), 3(e)).................................................................................................................................. 5

5. Vesica with proximal membranous portion forming a high and wide projection with a conspicuous pointed tip in profile, sclerotised portion without squamose ornamentation ( Figure 1 View Figure 1 (d)); juxta ovoid, in lateral view ( Figure 1 View Figure 1 (e))...................... T. (C.) pedunculata (Hall) View in CoL

5 ʹ. Vesica with proximal membranous portion forming a low and short projection with a bilobed tip in profile, sclerotised portion with squamose ornamentation ( Figure 3 View Figure 3 (e)); juxta not ovoid in lateral view, with narrower distal portion ( Figure 3 View Figure 3 (e))................................. ................................................................................................................................ T. (C.) alvarengai (Lopes) View in CoL

6. Mid femur with aposteroventral ctenidium; distiphallus apically globose; vesica without a proximal membranous portion ( Figure 8 (c,d); figs. 13, 14 in Lopes 1988).............................. .............................................................................................................................. T. (C.) ecuatoriana (Lopes) View in CoL

6 ʹ. Mid femur without a posteroventral ctenidium; distiphallus apically tapered; vesica with a short proximal membranous portion and sclerotised distal portion ( Figure 5 View Figure 5 (d,e)). ................................................................................................................................... T. (C.) luculenta (Lopes) View in CoL