Compsibidion Thomson, 1864

Santos-Silva, Antonio, Galileo, Maria Helena M. & Wappes, James E., 2017, Descriptions, notes and reassignments in Neoibidionini (Coleoptera: Cerambycidae: Cerambycinae) with a new genus, three new species and keys to species of Brechmoidion Martins, 1969, Compsibidion Thomson, 1864 and Rhysium Pascoe, 1866, Insecta Mundi 2017 (560), pp. 1-21 : 6-13

Key to species of Compsibidion Thomson, 1864 (adapted from Martins and Galileo 2007)

Obs. 1: For comparison images see Bezark (2017).

Obs. 2: Compsibidion trinidadense (Gilmour, 1963) was not included as it may not be a Compsibidion , but this cannot be determined without seeing the type.

1. Basal antennomeres not carinate dorsally (males unknown) .................................................... 2

– Basal antennomeres carinate dorsally at least from IV ............................................................. 5

2(1). Outer angle of elytral apex spiniform .......................................................................................... 3

– Outer angle of elytral apex unarmed ........................................................................................... 4

3(2). Antennomeres and legs yellow; elytral apex whitish; Nicaragua...... C. marqueti Audureau

– Antennomeres, legs and elytral apex dark brown; Brazil (Maranhão) ........................................ ................................................ C. maculatum Martins, Galileo and Limeira-de-Oliveira

4(2). Upper eye lobes with 5 rows of ommatidia; elytra at least partially with moderately abundant, decumbent pubescence; Bolivia........................................................ C. amboroensis sp. nov.

– Upper eye lobes with 3 rows of ommatidia; elytra with very sparse, decumbent pubescence; Bolivia................................................................................................. C. achiraensis sp. nov.

5(1). Upper eye lobes with 3 rows of ommatidia .................................................................................. 6

– Upper eye lobes with 4 or 5 rows of ommatidia (also see C. crassipede with 3 or 4 rows of ommatidia) ............................................................................................................................... 12

6(5). Prosternum without sericeous pubescence .................................................................................. 7

– Prosternum with sericeous pubescence ....................................................................................... 9

7(6). Elytral punctation restricted to poriferous punctures; Brazil (Amazonas, Pará) ....................... ............................................................................................................ C. sphaeriinum (Bates)

– Elytra with other, additional punctures between poriferous punctures .................................... 8

8(7). Center of pronotum sparsely pubescent; antennomere III in male cylindrical; antennae in female longer than body; Brazil (Minas Gerais, São Paulo) .......... C. megarthron (Martins, 1962)

– Center of pronotum without pubescence ; antennomere III in male fusiform; antennae in female only as long as body; Brazil (Bahia, São Paulo) ................... C. inornatum (Martins, 1962)

9(6). Outer angle of elytral apex unarmed; Brazil (Amazonas, Rondônia) .......................................... ................................................................................. C. uniforme Galileo and Martins, 2011

– Outer angle of elytral apex spiniform ........................................................................................ 10

10(9). Elytra mostly reddish brown, lacking distinct markings; Bolivia, Brazil (Maranhão) ............... ..................................................... C. ytu Martins, Galileo and Limeira-de-Oliveira, 2011

– Elytra with distinct markings .................................................................................................... 11

11(10). Elytra with transverse dark band at distal third; Panama......... C. mysticum Martins, 1969

– Elytra without transverse dark band at distal third; Brazil (Piauí) ............................................ ............................................................. C. pictum Galileo, Martins and Nascimento, 2014

12(5). Elytral setae whitish, thick ........................................................................................................ 13

– Elytral setae of other colors and/or slender ............................................................................... 14

13(12). Elytral setae more concentrated along suture, not organized in longitudinal rows in central area; prothorax equal in width at apex and base; Ecuador....................................................... ..................................................................................... C. decemmaculatum (Martins, 1960)

– Elytral setae equally distributed throughout, organized in rows in central area; prothorax wider at apex than base; Brazil (Goiás) .......................................... C. singulare (Gounelle, 1909)

14(12). Elytral color pattern simple, formed by light maculae and/or band on dark colors, or sometimes completely unicolorous ............................................................................................................ 29

– Elytral color pattern complex, formed by several bands or with moderately abundant contrasting punctures .................................................................................................................................. 15

15(14). Elytral apex lacking spine at outer angle .................................................................................. 16

– Elytral apex with spine at outer angle ...................................................................................... 17

16(15). Head dark brown; elytral apex with brown band; Colombia........................................................ ............................................................................... C. pumilium Martins and Galileo, 1999

– Head orangish; elytral apex without dark band; Brazil (Amazonas) ........................................... ........................................................................................................ C. aegrotum (Bates, 1870)

17(15). Elytra with sericeous pubescence very distinct on at least part, if not entire, surface .......... 18

– Elytra shining, pubescence sparse or absent ............................................................................ 20

18(17). Sericeous pubescence of elytra present only in distal half; French Guiana, Venezuela, Brazil (Amazonas, Pará) ................................................................. C. mulizonatum Martins, 1969

– Sericeous elytral pubescence present throughout ..................................................................... 19

19(18). Head black, with reddish macula on vertex; pronotum black, opaque, with reddish macula in center of base, center of basal region with whitish V-shaped pubescent area; Peru, Brazil (Pará, Mato Grosso, Goiás), Bolivia......................................... C. thoracicum (White, 1855)

– Head orangish, with dark bands; pronotum orangish, with dark anterior area, dark macula in central area, densely pubescent; Bolivia, Brazil (Goiás, Bahia, Minas Gerais, Espírito Santo, Rio de Janeiro, Paraná, Santa Catarina), Argentina (Misiones) .............................................. ................................................................................................... C. sommeri (Thomson, 1865)

20(17). Laterobasal tubercles of pronotum elevated and directed backward; antennal tubercles spiniform; Brazil (Bahia, Espírito Santo, Rio de Janeiro, São Paulo, Paraná, Santa Catarina, Rio Grande do Sul) ............................................................................................... C. zikani (Melzer, 1933)

– Laterobasal tubercles of pronotum not elevated and not directed backward; antennal tubercles rounded at apex ....................................................................................................................... 21

21(20). Basal antennomeres with long abundant setae; elytra yellowish, irregularly spotted by black or brown elevated punctures on base; Brazil (Amapá, Minas Gerais, Espírito Santo, Rio de Janeiro) ........................................................................................ C. amantei (Martins, 1960)

– Setae on basal antennomeres sparse and not especially long .................................................. 22

22(21). Basal quarter of elytra with black or brown macula/band ....................................................... 23

– Basal quarter of elytra without dark areas ............................................................................... 26

23(22). Elytra orangish, with 4 or 5 brown or reddish bands; Brazil (Pará) ............................................ ...................................................................................................... C. polyzonum (Bates, 1870)

– Elytra with 3 dark bands ............................................................................................................ 24

24(23). Each elytron with dark oblique fascia basally, narrowed from humerus to suture; Peru.......... ........................................................................................................ C. psydrum Martins, 1969

– Each elytron with distinct dark basal macula slightly widened toward suture ...................... 25

25(24). Club of meso- and metafemora entirely brown; Brazil (Espírito Santo, Rio de Janeiro) ............ ......................................................................................... C. guanabarinum (Martins, 1962)

– Club of meso- and metafemora partially darkened dorsally and ventrally; Ecuador, French Guiana, Brazil (Amazonas, Pará, Rondônia) .................................. C. charile (Bates, 1870)

26(22). Elytra with black, triangular area before midlength, delimited by whitish band anteriorly; Peru, French Guiana, Brazil (Amazonas, Pará) ......................................... C. rutha (White, 1855)

– Elytra with different color pattern ............................................................................................. 27

27(26). Elytra with orange, triangular area from base to middle, externally bordered by brown, narrow band; antennomeres III and IV in males brown and swollen, contrasting with the remaining antennomeres; French Guiana, Brazil (Amazonas, Pará) ............... C. basale (White, 1855)

– Elytra with different color pattern ............................................................................................. 28

28(27). Elytra pale yellow, with narrow, orange or orangish-brown bands; antennomere III not swollen in males; Mexico (Jalisco, Veracruz, Chiapas), Guatemala, El Salvador, Nicaragua, Costa Rica, Panama, Venezuela, Guyana, French Guiana, Brazil (Amazonas, Pará, Mato Grosso, Goiás, Maranhão, Alagoas, Ceará, Bahia, Minas Gerais, Espírito Santo, Rio de Janeiro, São Paulo, Paraná, Santa Catarina, Rio Grande do Sul), Argentina (Misiones) ............................ ...................................................................................................... C. vanum (Thomson, 1867)

– Elytra pale yellow, spotted by black maculaeand punctures; antennomere III in males thickened; Brazil (Espírito Santo) .................................................. C. balium Napp and Martins, 1985

29(14). Elytral apex unarmed ................................................................................................................. 30

– Elytral apex spiniform ................................................................................................................ 42

30(29). Entire distal half of elytra dark, contrasting with lighter-colored basal half; Bolivia................ .................................................................................................. C. reichardti (Martins, 1962)

– Distal half of elytra partially, or not at all, contrasting in color with basal half .................... 31

31(30). Elytra without contrasting maculae and/or bands; Brazil (Rio de Janeiro, Paraná, Santa Catarina) ....................................................................................... C. niveum (Martins, 1962)

– Elytra with contrasting maculae and/or bands ......................................................................... 32

32(31). Each elytron with a single pale yellow macula before middle, distinctly removed from suture; Ecuador................................................................... C. in fl atum Martins and Galileo, 2014

– Each elytron without or with two, or more pale yellow maculae (if only a single macula, it is horizontal, placed at about midlength and reaching or nearly reaching suture) ................ 33

33(32). Each elytron with large triangular yellow macula near middle; Colombia................................. ................................................................................................... C. paradoxum Martins, 1971

– Each elytron without large triangular yellow macula near middle ......................................... 34

34(33). Apical quarter of elytra with distinct, well delimited pale yellow macula .............................. 35

– Apical quarter of elytra without pale yellow macula ................................................................ 36

35(34). Basal 2/3 of each elytron with a large, pale yellow, elliptical macula clearly removed from base; Brazil (Minas Gerais) ......................................... C. novalimae Martins and Galileo, 2012

– Basal 2/3 of each elytron with a large, pale yellow macula almost reaching base, partially and obliquely divided by brown band; Brazil (Bahia, Minas Gerais, Espírito Santo) .................... .................................................................................................... C. capixaba (Martins, 1962)

36(34). Pale yellow band of elytra placed near center, horizontal (not slanted) .................................. 37

– Pale yellow band of elytra placed near center, ascending from side to suture ........................ 38

37(36). Antennomeres III and IV in males widened, carinate, and without abrupt change in thickness compared to remaining antennomeres; Bolivia, Brazil (Goiás, Bahia, Minas Gerais, Espírito Santo, Rio de Janeiro, São Paulo, Paraná, Santa Catarina, Rio Grande do Sul), Paraguay, Argentina (Misiones) ........................................................... C. truncatum (Thomson, 1865)

– Antennomeres III and IV in males with abrupt change in thickness compared to remaining antennomeres, slightly carinate only at base; Venezuela.............. C. clivum Martins, 1971

38(36). Pale yellow maculae of elytra not well delimited or not distinctly contrasting with surrounding surface, or completely absent .................................................................................................. 39

– Pale yellow maculae of elytra very conspicuous, well delimited and contrasting with remaining surface ...................................................................................................................................... 40

39(38). Metafemora in male notably widened; elytra slightly longer than twice length of prothorax; Brazil (Bahia, São Paulo, Mato Grosso do Sul), Paraguay, Argentina (Salta) ......................... ..................................................................................................... C. crassipede Martins, 1971

– Metafemora in male not distinctly widened; elytra about 3 times longer than prothorax; Brazil (Minas Gerais, São Paulo) ........................................... C. triviale Napp and Martins, 1985

40(38). Apical elytral pale yellow macula V-shaped; Uruguay.................... C. monnei Martins, 1969

– Apical elytral pale yellow macula not V-shaped, usually oblique, J-shaped or U-shaped ...... 41

41(40). Antennomeres III and IV in male not tumid; apical pale yellow macula of each elytron narrower and more elongate than basal macula; Bolivia, Brazil (Bahia), Argentina (Salta, Catamarca, Tucumán, Santiago del Estero, La Rioja, Córdoba, San Juan, Chaco, Entre Ríos, Buenos Aires), Paraguay................................................................... C. circun fl exum Martins, 1971

– Antennomeres III and IV in male tumid; apical pale yellow macula of each elytron about as wide as basal macula; Bolivia, Brazil (Maranhão, Goiás, Mato Grosso, Sergipe, Alagoas, Bahia, Minas Gerais, Espírito Santo, Rio de Janeiro, São Paulo, Paraná, Santa Catarina, Rio Grande do Sul), Paraguay, Argentina (Jujuy, Salta, Catamarca, Tucumán, Santiago del Estero, Chaco, Buenos Aires), Uruguay...................................................... C. fairmairei (Thomson, 1865)

42(29). Elytra with a transverse yellowish-white band at midlength .................................................. 43

– Elytra with different markings .................................................................................................. 45

43(42). Antennal tubercles distinctly projected and acute at apex; antennomeres III–VI in male slightly widened, without abrupt change in thickness compared to remaining antennomeres; Brazil (Bahia, Minas Gerais, Rio de Janeiro) .......................................... C. divisum Martins, 1969

– Antennal tubercles slightly projected, with apices nearly blunt; antennomere III in male tumid, with abrupt change in thickness compared to IV .................................................................. 44

44(43). Apices of femora acutely spiny; humeri reddish; antennomere III not carinate in male; Brazil (Goiás, Espírito Santo, São Paulo) ................................. C. unifasciatum (Gounelle, 1909)

– Apices of femora with triangular lobes ; entire anterior half of elytra reddish; antennomere III carinate in male; Brazil (Espírito Santo) ................................... C. carenatum Martins, 1969

45(42). Elytra unicolorous except for dark apex .................................................................................... 46

– Elytra with light maculae and/or bands .................................................................................... 47

46(45). Elytra, scape, basal antennomeres and legs pale yellow; Brazil (Bahia, Minas Gerais, Rio de Janeiro) ....................................................................... C. nigroterminatum (Martins, 1965)

– Elytra reddish; scape, basal antennomeres and legs dark brown or black; Brazil (Bahia, Minas Gerais, Rio de Janeiro) ........................................................ C. trichocerum (Martins, 1962)

47(45). Distal area of elytra same color as base, without light bands and/or maculae ....................... 48

– Distal area of elytra with band or macula, or distinctly contrasting with basal color ............ 66

48(47). Pronotal pubescence not arranged in bands .............................................................................. 49

– Pronotal pubescence arranged into either two longitudinal bands or a V-shaped pattern .... 57

49(48). Elytra dark, with light macula and/or fascia ............................................................................ 50

– Elytra with different color pattern (general color usually reddish) ......................................... 52 50(49) Apices of meso- and metafemora not spiny; Bolivia...................................................................... ........................................................................... C. morrisi Galileo and Santos-Silva, 2016

– Apices of meso- and metafemora distinctly spiny, longer on inner side .................................. 51

51(50) General appearance robust; antennomeres III–VI in male distinctly tumid; spines at apices of meso- and metafemora short; Brazil (Santa Catarina) ........ C. meridionale Martins, 1969

– General appearance slender ; antennomeres III–VI in male slightly tumid; spines at apices of meso- and metafemora long; Brazil (Maranhão, Bahia, Minas Gerais, Espírito Santo, Rio de Janeiro, São Paulo, Paraná, Santa Catarina) ....................... C. cleophile (Thomson, 1865)

52(49). Base and apex of elytra with same color ................................................................................... 53

– Base and apex of elytra with different colors ............................................................................ 55

53(52). Each elytron with one large pale yellow band from near base to past midlength; Brazil (Goiás, Piauí) ......................................................................... C. elianae Martins and Galileo, 2012

– Each elytron with two pale yellow maculae .............................................................................. 54

54(53). Each elytron with two elliptical or subrounded pale yellow maculae; Bolivia, Brazil (Mato Grosso) .......................................................................... C. ybyra Martins and Galileo, 2012

– Each elytron with pale yellow band in basal half and another wide, oblique one ascending from margin to suture in apical half; Brazil (Mato Grosso) ......... C. derivativum Martins, 1971

55(52). Prothorax reddish; basal pale yellow macula of elytron small and rounded; Brazil (Espírito Santo) ......................................................................................... C. tuberosum Martins, 1971

– Prothorax dark; basal pale yellow macula of elytron narrow and elongate ............................ 56

56(55). Metafemora mostly dark brown; antennomere III in male not tumid; Brazil (São Paulo) .......... ...................................................................................................... C. paulista (Martins, 1962)

– Metafemora light reddish; antennomere III in male tumid; Brazil (Minas Gerais) ................... ........................................................................................... C. peti Martins and Galileo, 2012

57(48). Pronotal pubescence forming V-shaped pattern ....................................................................... 58

– Pronotal pubescence arranged into two longitudinal bands ..................................................... 61

58(57). Pronotal pubescence confined to base and inner side of laterobasal tubercles; Brazil (Bahia, Minas Gerais, Espírito Santo) .................................. C. concisum Napp and Martins, 1985

– Pronotal pubescence covers laterobasal tubercles and also reaches laterodistal tubercles .... 59

59(58). Antennomere III in male tumid and not carinate; light macula in basal half of elytron elliptical; Brazil (Espírito Santo) ................................................ C. punga Martins and Galileo, 2007

– Antennomere III in male tumid and carinate; light macula in basal half of elytron distinctly longitudinal .............................................................................................................................. 60

60(59). Prothorax reddish; elytron lacking pale yellow macula close to margin in basal half; Brazil (Rio Grande do Sul) ........................................................................... C. rubricolle (Melzer, 1935)

– Prothorax dark brown; elytron with pale yellow macula close to margin in basal half; Ecuador, Bolivia, Brazil (Pará, Goiás, Mato Grosso, Maranhão, Bahia, Minas Gerais, Espírito Santo, Rio de Janeiro, São Paulo, Paraná, Santa Catarina, Rio Grande do Sul), Paraguay, Argentina (Jujuy, Salta, Tucumán, Misiones, Corrientes) ................. C. graphicum (Thomson, 1867)

61(57). Elytra with basal half reddish and apical half black, separated by pale yellow band which projects forward along suture; Brazil (Alagoas, Bahia, Minas Gerais, Espírito Santo, Rio de Janeiro) ..................................................................................... C. simillimum Martins, 1969

– Basal and apical halves of elytra with same color .................................................................... 62 62(61). Elytra reddish, with pale yellow maculae ................................................................................. 63

– Elytra dark, with pale yellow maculae ...................................................................................... 64

63(62). Distal lobes of meso- and metafemora acute; antennomeres III and IV in males thickened; Venezuela, Peru, Bolivia, Brazil (Mato Grosso, Goiás, Bahia, Minas Gerais, Espírito Santo, Rio de Janeiro, São Paulo, Santa Catarina) ................................. C. ilium (Thomson, 1864)

– Inner distal lobe of mesofemora spined ; both lobes of metafemora spined (inner longer); antennomeres in males not thickened; Brazil (Goiás, Bahia, Minas Gerais, Espírito Santo, Rio de Janeiro, São Paulo, Paraná, Santa Catarina) .... C. quadrisignatum (Thomson, 1865)

64(62). Anterolateral tubercles of pronotum rounded, of moderate size; light band in middle of elytron narrow, oblique, ascending from margin to suture; Brazil (Pernambuco, Bahia, Minas Gerais, Espírito Santo, Rio de Janeiro) .............................................. C. angulare (Thomson, 1867)

– Anterolateral tubercles of pronotum subacute, distinct; light band in middle of elytron completely or partially transverse ............................................................................................................. 65

65(64). Pale macula and band wide, covering more than half of elytra; larger species (length 13.7–16.3 mm); antennomere III in male carinate; Venezuela, Bolivia, French Guiana, Brazil (Pará, Piauí) ......................................................................................... C. callispilum (Bates, 1870)

– Pale macula and band narrow, covering less than one-fourth of elytra, smaller species (length 8.2 mm); antennomere III in male slightly carinate; Brazil (Mato Grosso) ............................. ............................................................................................ C. melancholicum Martins, 1969

66(47). Pronotal pubescence covering entire surface except tip of central tubercle; if covering lesser part of surface, then not arranged in bands or V-shaped ...................................................... 67

– Pronotal pubescence arranged in two bands, V-shaped pattern, or subparallel (reaching at most anterolateral tubercles) ........................................................................................................... 70

67(66). Basal half of elytra pale yellow, slightly reddish only at base, with light macula of each elytron elongate; light apical band of each elytron placed at apical third; antennomeres III and IV in male thickened; Brazil (Espírito Santo) ................................. C. muricatum Martins, 1971

– Basal half of elytra from reddish to black ................................................................................. 68

68(67). Pale yellow band of elytron nearly horizontal; Peru, Ecuador, French Guyana, French Guiana, Brazil (Amazonas, Pará) ............................................................... C. tethys (Thomson, 1867)

– Pale yellow band of elytron ascending from margin to suture ................................................. 69

69(68). Antennomeres III and IV in male not thickened; distal lobes of meso- and metafemora rounded; Panama, Trinidad and Tobago, Venezuela, Colombia, Guyana, Suriname, French Guiana, Peru, Bolivia, Brazil (Amapá, Amazonas, Pará, Rondônia, Mato Grosso, Maranhão) ............ ............................................................................................. C. maronicum (Thomson, 1867)

– Antennomeres III and IV in male thickened; inner distal lobe of mesofemur projected, distal lobes of metafemora spiniform; Brazil (Piauí) ........................................................................... .................................................................... C. paragraphycum Martins and Galileo, 2013

70(66). Pronotal pubescence V-shaped ................................................................................................... 71

– Pronotal pubescence not V-shaped ............................................................................................. 74

71(70). Elytra lacking narrow, subcircular dark bands in basal half; Brazil (Amazonas). ..................... .............................................................................. C. manauara Martins and Galileo, 2012

– Elytra with narrow, subcircular dark bands in basal half ....................................................... 72 72(71). Elytron with narrow, elongate, longitudinal pale yellow band in basal half; Ecuador, Bolivia, Brazil (Pará, Goiás, Mato Grosso, Maranhão, Bahia, Minas Gerais, Espírito Santo, Rio de Janeiro, São Paulo, Paraná, Santa Catarina, Rio Grande do Sul), Paraguay, Argentina (Jujuy, Salta, Tucumán, Misiones, Corrientes) ............................. C. graphicum (Thomson, 1867)

– Elytron without pale yellow longitudinal band in basal half ................................................... 73

73(72). Antennomeres III and IV not thickened in male; larger species; Brazil (Pará) .......................... ................................................................................................................ C. orpa (White, 1855)

– Antennomeres III and IV thickened in male; smaller species; Brazil (Pará) .............................. ..................................................................................... C. taperu Martins and Galileo, 2007

74(70). Elytra with basal half reddish or yellowish and apical half black (except apex) .................... 75

– Elytra unicolorous (except apex) ................................................................................................ 76

75(74). Elytron without light macula in basalhalf; pubescence of pronotum not surpassing anterolateral tubercles; Peru............................................................................. C. virgatum Martins, 1969

– Elytron with light macula in basal half; pubescence of pronotum surpassing anterolateral tubercles; Bolivia, Brazil (Maranhão, Piauí, Alagoas, Paraíba, Bahia, Mato Grosso, Mato Grosso do Sul, Goiás, Minas Gerais, São Paulo), Argentina (Jujuy, Salta, Tucumán, Santiago del Estero) ............................................................................. C. campestre (Gounelle, 1909)

76(74). Elytral base with reddish area; Panama, Colombia, Venezuela.................................................. .................................................................................................. C. litturatum (Martins, 1960)

– Elytral base without reddish area ............................................................................................. 77

77(76). Light macula in basal half of elytron reaching humerus; Colombia, Venezuela......................... ..................................................................................................... C. varipenne Martins, 1969

– Light macula in basal half of elytron not reaching humerus ................................................... 78

78(77). Central band of elytron curved, not reaching margin or suture; Brazil (Goiás) ......................... ........................................................................................................ C. omissum Martins, 1969

– Central band of elytron straight ................................................................................................ 79

79(78). Elytron with small, rounded, pale subapical macula (not covering apex), and a narrow central band (about as wide as antennomere III); Brazil (Pernambuco, Bahia, Minas Gerais, Espírito Santo, Rio de Janeiro) ............................................................. C. angulare (Thomson, 1867)

– Elytron with pale apical macula extending on apex proper, and a wide central band (about twice width of antennomere III) ....................................................................................................... 80

80(79). Central oblique band of elytron narrowly separated from basal macula (distance between them nearly equal to width of antennomere III); Brazil (Goiás, Mato Grosso, Piauí) ...................... ............................................................................................... C. decoratum (Gounelle, 1909)

– Central oblique band of elytron widelyseparated from basal macula (distance between them more than 3 times width of antennomere III); Bolivia, Brazil (Goiás, Mato Grosso, Mato Grosso do Sul, Maranhão, Piauí, Alagoas, Paraíba, Bahia, Minas Gerais, São Paulo), Argentina (Jujuy, Salta, Tucumán, Santiago del Estero) ................................. C. campestre (Gounelle, 1909)