Compsibidion achiraensis, Santos-Silva & Galileo & Wappes, 2017

Santos-Silva, Antonio, Galileo, Maria Helena M. & Wappes, James E., 2017, Descriptions, notes and reassignments in Neoibidionini (Coleoptera: Cerambycidae: Cerambycinae) with a new genus, three new species and keys to species of Brechmoidion Martins, 1969, Compsibidion Thomson, 1864 and Rhysium Pascoe, 1866, Insecta Mundi 2017 (560), pp. 1-21 : 4-5

Compsibidion achiraensis sp. nov.

( Fig. 7–11 , 18 )

Description. Holotype female. Head and prothorax mostly dark brown dorsally, gradually reddish brown toward ventral side; mouthparts mostly yellowish brown; antennae reddish brown; ventral side of meso- and metathorax and abdominal ventrites mostly reddish brown, lighter or darker in some areas; legs dark reddish brown; elytra dark brown except for irregular, wide yellow spot near midlength. Pubescence and erect setae yellowish white.

Head. Frons finely, densely, confluently punctate, laterally separated from antennal tubercles by oblique, deep depression; pubescence not obscuring integument. Area between antennaltubercles finely, confluently punctate, with narrow carina on each side of coronal suture; pubescence not obscuring integument. Area between upper eye lobes with slightly elevated, subelliptical plate centrally (depressed toward posterior ocular edge, surpassing posterior ocular edge), finely, confluently punctate (punctures coarser than on frons); pubescence not obscuring integument. Remaining surface of vertex finely, abundantly punctate; pubescence not obscuring integument; with one long, erect seta on each side near eye. Area behind upper eye lobes finely striate-punctate close to eye, finely, sparsely punctate toward prothorax; pubescence clearly not obscuring integument close to eye, glabrous toward prothorax. Area behind lower eye lobes moderately coarsely, sparsely punctate close to eyes, finely, sparsely punctate toward prothorax; with long, erect, sparse setae close to eyes, glabrous toward prothorax. Antennal tubercles finely, abundantly punctate; pubescence not obscuring integument, slightly denser than on frons. Genae finely, abundantly punctate, except smooth area close to apex; with sparse pubescence and some long, erect setae. Postclypeus glabrous, smooth laterally, narrow finely, abundantly punctate centrally; with one long, erect seta on each side close to glabrous area. Labrum glabrous, smooth, coplanar with anteclypeus except inclined in center of distal area, fringed with setae. Distance between upper eye lobes 0.45 times length of scape; distance between lower eye lobes 0.65 times length of scape; upper eye lobes with 3 rows of ommatidia. Antennae 1.75 times elytral length, reaching elytral apex at apex of antennomere IX. Scape subcylindrical, slightly widened toward apex, arched in side view; with some long, erect setae throughout. Antennomeres not carinate. Antennomeres III–V with long, erect, sparse setae ventrally (sparser toward V). Antennal formula (ratio) based on length of antennomere III: scape = 0.75; pedicel = 0.19; IV = 0.77; V = 0.85; VI = 0.87; VII = 0.87; VIII = 0.81; IX = 0.75; X = 0.67; XI = 0.67.

Thorax. Prothorax subcylindrical, 1.35 times longer than wide, with similar basal and distal constrictions. Pronotum with five slightly elevated gibbosities (one on each side of basal quarter; one on each side of anterior third; one centrally at about midlength); moderately coarsely, very sparsely punctate; somewhat opaque, microsculptured, except large, smooth, shining area medially (this area including anterolateral and central gibbosities); pubescence partially obscuring integument, interspersed with long, erect setae, except in glabrous smooth area. Sides of prothorax mostly smooth, except striate basal and distal areas (more distinct in basal area); nearly glabrous centrally, with short, sparse setae toward pronotum and ventral side; with a few long, erect setae in anterior third. Prosternum with short, sparse setae throughout, more abundant in basal half (forming slightly distinct V-shaped pubescent area); nearly smooth in basal half, finely, transversely striate in distal half. Prosternal process narrowly constricted centrally. Mesosternum sparsely pubescent centrally, denser laterally. Metepisternum and metasternum with pubescence moderately abundant, but not obscuring integument. Scutellum with pubescence not obscuring integument.

Elytra. Finely, sparsely punctate, interspersed with coarse punctures; with minute, sparse, decumbent setae throughout, interspersed with long, erect, sparse setae (slightly denser in distal third); sides slightly widened behind basal half; apex rounded.

Legs. Femora pedunculate-clavate (mainly meso- and metafemora); pubescence not obscuring integument, interspersed with long, erect, sparse setae. Tibiae not longitudinally carinate. Metatarsomere Iabout as long as II–III together.

Abdomen. Ventrites with abundant pubescence, not obscuring integument, interspersed with long, erect setae; apex of ventrite V rounded.

Dimensions (holotype female). Total length 8.25; prothorax: length 1.45; anterior width 1.15; posterior width 1.10; max. prothoracic width 1.25; humeral width 1.70; elytral length 5.65.

Type material. Holotype female from BOLIVIA, Santa Cruz: Achira Campo , 9-11.X.2004, Wappes & Morris col. ( MNKM).

Etymology. Named after Achira Campo, where the type was collected.

Remarks. The elytral pattern of Compsibidion achiraensis sp. nov. is similar to that of many species placed in several genera of Neoibidionini . In Compsibidion it is most similar to C. inflatum Martins and Galileo, 2014 and C. amboroensis sp. nov. It differs from the former primarily by the prothorax more cylindrical, antennomere IV slightly shorter than V, and elytra with sparse long, erect setae. In females of C. inflatum the prothoracic constrictions are also stronger, making the central area wider, the antennomere IV is distinctly shorter than V, and the elytra clearly have more abundant erect setae. It differs from C. amboroensis as follows: ommatidia slightly coarser; upper eye lobes with 3 rows of ommatidia ( Fig. 18 ); prothorax longer and narrower; and decumbent elytral setae sparser throughout. In C. amboroensis the ommatidia are finer, upper eye lobes with 5 rows of ommatidia ( Fig. 17 ), prothorax shorter and wider, and decumbent elytral setae distinctly denser toward apex.

Figures

Figures 1–11. Newspecies of Neoibidionini.1–6, Chiquitanovolcanesensis sp. nov. 1) Holotype male, dorsal habitus. 2) Holotype male, ventral habitus. 3) Holotype male, lateral habitus. 4) Holotype male, head, frontal view. 5) Holotype male, head and scape, dorsal view. 6) Paratype female, dorsal habitus. 7–11, Compsibidion achiraensis sp. nov., holotype female: 7) Head, frontal view. 8) Head and scape, dorsal view. 9) Dorsal habitus. 10) Ventral habitus. 11) Lateral habitus.

Figures 12–18. New species of Compsibidion. 12–17, Compsibidion amboroensis sp. nov. 12) Dorsal habitus. 13) Ventral habitus. 14) Lateral habitus. 15) Head, frontal view. 16) Head and scape, dorsal view. 17) Eye, frontal view. 18, Compsibidion achiraensis sp. nov.: holotype female, eye, frontal view.