Endroedyolini, Davis, Deschodt & Scholtz, 2019

Daniel, Gimo M., Strümpher, Werner P. & Deschodt, Christian M., 2022, An unexpected new flightless dung beetle species (Coleoptera: Scarabaeidae: Scarabaeinae: Endroedyolini) from the Cederberg Mountains, South Africa, Zootaxa 5093 (4), pp. 414-420 : 419

Identification key to Endroedyolini , adapted and modified from Deschodt et al. (2020).

1. Metaventrite anteriorly with deep fovea on each side of midline, pronotal disc on each side near posterior angle with short elevated carina ................................................ Endroedyolus paradoxus Scholtz & Howden, 1987

- Metaventrite anteriorly without deep fovea on each side of midline, and pronotal disc with or without short elevated carina on each side near posterior angle........................................................................... 2

2. Pronotal disc with short elevated carina on each side near posterior angle................................................................................................... Aliuscanthoniola similaris Deschodt & Scholtz, 2008

- Pronotal disc without short, elevated carina on each side near posterior angle ...................................... 3

3. Mesoventrite medially wide, or wide enough for, more than three irregular rows of punctures......................... 4

- Mesoventrite medially narrow, with at most two irregular rows of punctures (punctures sometimes absent in this area)..... 5

4. Mesometaventral suture angulate at anterior apex.................. Outenikwanus tomentosus Scholtz & Howden, 1987

- Mesometaventral suture rounded at anterior apex...................... Upsa centennial Deschodt, Sole & Scholtz, 2020

5. Elytron in lateral view with depression on lateral side ......................................................... 6

- Elytron in lateral view without depression on posterolateral side ................................................. 7

6. Pronotal depression on mid-basal edge present, mesometaventral suture rounded; lateral elytral spine in males present.................................................................. Silvaphilus oubosiensis Roets & Oberlander 2010

- Pronotal depression on mid-basal edge absent, mesometaventral suture curving very slightly anteriorly; lateral elytral spine in males absent ....................................... Silvaphilus joselmae Daniel, Strümpher & Deschodt new species

7. Mesometaventral suture rounded at anterior apex............................................................ 8

- Mesometaventral suture angulate at anterior apex........................................................... 10

8. Protibial teeth 2 and 3 small, almost half the size of protibial tooth 1........... Peckolus parvus Scholtz & Howden, 1987

- Protibial teeth 2 and 3 minute, much less than half the size of protibial tooth 1..................................... 9

9. The recurved setae on the body almost half the length of the straight setae...... Peckolus alpinus Howden & Scholtz, 1988

- The recurved setae on the body barely discernible..................... Peckolus poenskopius Deschodt & Scholtz, 2008

10. Lateral part of mesometaventral suture (between mesocoxae) converging anteriorly ........................................................................................... Parvuhowdenius harrisoni Deschodt & Scholtz, 2008

- Lateral part of mesometaventral suture (between mesocoxae) parallel........................................... 11

11. Punctures on mesoventrite large and clear, separated by less than one puncture diameter.......................................................................................... Nebulasilvius insularis Deschodt & Scholtz, 2008

- Punctures on mesoventrite small and separated by at least one puncture diameter .................................................................................................. Nebulasilvius johani Deschodt & Scholtz, 2008