Tetana, Brunner von Wattenwyl, 1878
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publication ID |
https://doi.org/ 10.11646/zootaxa.2944.1.1 |
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persistent identifier |
https://treatment.plazi.org/id/03B087E5-5B4A-FFA3-FF79-F6E2FAD4FD8E |
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treatment provided by |
Felipe |
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scientific name |
Tetana |
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Tetana View in CoL group
Ten species are included in the Tetana group. These are generally very darkly colored species, dorsum piceous, except two species that have a pronotal fascia ( H. biundulata and H. penultimata ) ( Figs. 66 View FIGURE 66 , 67 View FIGURE 67 , 70 View FIGURE 70 , 71 View FIGURE 71 , 74 View FIGURE 74 , 75 View FIGURE 75 , 78 View FIGURE 78 , 79 View FIGURE 79 , 82 View FIGURE 82 , 83 View FIGURE 83 ). Males of four of the species are immediately recognized by a prominent midventral tubercle on the first abdominal ventrite (e.g., Fig. 305 View FIGURES 304–309 ); these species are H. tetana , H. insandalia , H. acumena , and H. aulaarta . Males of three other species have a different abdominal sexual dimorphism: the fifth ventrite is raised in a transverse ridge, abruptly declivous behind; species with this character are H. tarsotricha , H. tricosipes , and H. huonica . The meso- and metatibiae are not markedly modified, but seven of the species have long hair-like setae on the tarsi of those legs (e.g., Fig. 314 View FIGURES 310–315 ).
Most of the species have very large plaques that are very narrowly separated, more narrowly separated posteriorly than anteriorly, which is unusual in the genus (e.g., Figs. 308, 309 View FIGURES 304–309 , 311 View FIGURES 310–315 ). The pronotal foveae are reduced in all but two species: PF1 and PF4 are absent, and PF2 is extremely shallow, obsolete. Several of the species are very similar in most characters, and will require examination of the male genitalia for reliable determinations.
A characteristic feature of the aedeagus in this group is the shape of the left paramere, which is usually moreor-less oval in cross-section, very strong and stiff, and arches up and over the distal piece (e.g., Figs. 68, 69). Two species, H. penultimata and H. altapapua , have plaques shaped differently than the other species ( Figs. 78 View FIGURE 78 , 83 View FIGURE 83 ), and also have quite distinct male genitalia ( Figs. 80 View FIGURES 80–81 , 85 View FIGURES 84–85 ); these two species, however, have the characteristic arching left paramere and are therefore provisionally placed in the Tetana group. The female tergite X, gonocoxite, and spermatheca of H. tetana , H. tarsotricha , and H. biundulata are illustrated ( Figs. 395–397); they are quite similar, with (as in the male genitalia) H. biundulata being slightly different from the other two species.
Members of this group have been collected at an elevation range of 500–2680 m. Seven of the ten species have not been collected above 1700 m; the three other species have been collected at a range of 970–2680 m (maps Figs. 459–468 View FIGURES 459–462 View FIGURES 463–466 View FIGURES 467–470 ). The distributions suggest that this is primarily an Area 1 group, with some species spreading to the Finisterre Mts. (Area 5) and the Herzog Mts. (Area 6); only one of the ten species in the group, H. huonica , is endemic to Area 5, and no species is endemic to Area 6. Most microhabitat data indicate gravel banks of streams or rivers, with a few pools or ponds mentioned. Representative microhabitat notations include the following: washed from margins of riverside still water pools in rainforest, gravel and cobble dominated microhabitats, but some muddy; muddy gravel banks of small shaded stream; gravel bank of small river in places where rotten leaves had accumulated; washed from margins of small pool of a steep, rocky, forest stream at confluence with river, gravel, cobbles, wet rock microhabitats; gravel banks of small clear river in dense forest; montane stream tumbled down among moss-covered rocks which alternated with short sandy runs in more level sections, almost completely shaded.
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