Hydraena fundacta, Perkins, 2011

Perkins, Philip D., 2011, New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944, Zootaxa 2944 (1), pp. 1-417 : 141

Hydraena fundacta , new species

( Figs. 262 , 264 , 552 )

Type Material. Holotype (male): Madang Province: Adelbert Mts. , Sewan –Keki, 700 m, 4° 42.215' S, 145° 25.154' E, 4 v 2006, Balke & Manaono ( PNG 51) ( ZSM) . Paratypes (32): Madang Province: Same data as holotype (20 MCZ, NHM, NMW, PNG, ZSM) ; Adelbert Mts. , below Keki, 790 m, 4° 42.3' S, 145° 25.089' E, 5 v 2006, Balke & Manaono ( PNG 53) (2 ZSM) ; Sandaun Province: Bewani Stn. , limestone creek at base of Bewani Mts., 400 m, 3° 5.13' S, 141° 10.227' E, 12 iv 2006, Balke & Sagata ( PNG 39) (3 ZSM) ; road Vanimo–Bewani close coast, 400–480 m, 2° 44.372' S, 141° 15.067' E, 13 iv 2006, Balke & Sagata ( PNG 41) (7 MCZ, NHM, NMW, PNG, ZSM) .

Differential Diagnosis. The putative sister species of H. johncoltranei ( Figs. 262 , 263 ); refer to the differential diagnosis of that species.

Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.30/0.57; head 0.18/0.32; pronotum 0.33/0.49, PA 0.37, PB 0.43; elytra 0.77/0.57. Dorsum of head dark brown, clypeus and labrum slightly lighter; pronotum testaceous around dark brown, diffusely margined, rectangular macula, ratios of color bands, as measured in midline, ca. 7/12/5; elytra brown; legs light brown; maxillary palpi testaceous, tip not darker. Dorsum moderately shining.

Frons punctures ca. 1xef near eyes, smaller and slightly sparser medially; interstices weakly shining, 1–2xpd. Clypeus effacedly microreticulate laterally, very finely sparsely punctulate and weakly shining medially. Mentum very sparsely very finely punctulate, shining. Postmentum effacedly microreticulate in shallow median depression, otherwise finely sparsely punctate. Genae very slightly raised, shining, with shallow, obsolete median impression, without posterior ridge. Pronotum transverse, ca. median 3/4 of anterior margin slightly arcuate to posterior, sides weakly arcuate, slightly emarginate between midlength and posterior angle; punctures on disc ca. 1xpd largest frons punctures, interstices shining, ca. 2–3xpd, punctures anteriorly and posteriorly slightly larger than those on disc; PF1, PF2 and PF4 absent; PF3 shallow, wide.

Elytra widest slightly anterior to midlength; summit of posterior declivity at or very near midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, a few punctures subserial, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 2–3xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming angle with one another.

P1 slightly wider than laminate; median carina nearly straight in profile, slightly arcuate between coxae. P2 markedly concave, length/width ca. equal, sides parallel, apex blunt, not concave. Plaques absent. Metaventrite flat or very slightly convex between mesocoxae. AIS width at straight posterior margin slightly greater than P2. All legs of moderate length. Profemur (male) with very small tubercle on ventral margin near basal 1/3; protibia very slightly arcuate, width quite uniform in distal 3/4. Meso- and metatibia straight, slender. Abdominal apex (male) enlarged, asymmetrical; last tergite with oblique ridge on dorsal surface. Aedeagus as illustrated ( Fig. 264 ).

Etymology. Named in reference to the geographical distribution near the ocean, and membership in the Fundarca group.

Distribution. The only species exclusive shared between Areas 4 and 5; elevation range 400–790 m ( Fig. 552 ).

Figures

FIGURE 262. Hydraena fundacta, holotype, dorsal and lateral habitus, and venter.

FIGURE 263. Hydraena johncoltranei, holotype, dorsal and lateral habitus, and venter.

FIGURES 264–265. Aedeagi of holotypes. 264. H. fundacta. 265. H. johncoltranei.

FIGURES 551–554. Geographical distributions of Hydraena species.