Carya cf. minor SAPORTA , 1873

Denk, Thomas, Sami, Marco, Teodoridis, Vasilis & Martinetto, Edoardo, 2022, The Late Early Pleistocene Flora Of Oriolo, Faenza (Italy): Assembly Of The Modern Forest Biome, Fossil Imprint 78 (1), pp. 217-262 : 234

Carya cf. minor SAPORTA, 1873

Text-fig. 9a–e

1873 Carya minor SAPORTA , p. 217.

1876 Juglans minor (SAPORTA) SAPORTA et MARION , p. 166, pl. 37, figs 1–6.

1970 Carya cf. minor SAPORTA ; Pavia, pl. 3, fig. 5.

2001 Carya cf. minor SAPORTA ; Martinetto and Sami, p. 17, textfig. 10.

M a t e r i a l. Oriolo MSF 631, 631-1, 631-2, 634, 634-1, 635, 639, 690, 691, 755, 756, 757-1, 758, 759, 760, 761, 762, 763, 764, 765, 766, 767, 768, 769, 770, 771, 772, 773, 774, 775, 777.

D e s c r i p t i o n. Leaflet, petiolate, petiole <2 mm long, lamina elliptic, ovate or obovate, 39–124 mm long, 18–57 mm wide, base cuneate decurrent, acute rounded, apex acute or acuminate, secondary venation mixed brochidodromous, semicraspedodromous, secondary veins branching close to leaf margin or rarely close to primary vein, secondary veins and intersecondary veins spaced 2–4 per 1 cm primary vein, margin sharply serrate.

R e m a r k s. The material from Oriolo is closely similar to leaves/leaflets of C. minor from the Pliocene of Meximieux ( France) and Willershausen ( Germany) in some instances (densely spaced, regular secondaries forming a brochidodromous pattern of loops), while other leaflets have a more irregular pattern of secondary veins. From Miocene strata of France and Crete, Kvaček et al. (2011) and Zidianakis et al. (2020) described dispersed leaves and leaflets of Carya as Carya sp. , accounting for differences in secondary venation with known western Eurasian Neogene species of Carya . Pavia (1970) reported a very similar leaflet from Arboschio (Pliocene).

Figures

Text-fig. 9. a–e: Carya aff. minor. a: Terminal leaflet, Oriolo MSF 755. b: Oriolo MSF 755. c: Oriolo MSF 766. d: Oriolo MSF 768. e: Oriolo MSF 635. f, g: Pterocarya aff. fraxinifolia. f: Fruit, Oriolo MSF 632. g: Leaflet, Oriolo MSF 632a. h–j: Alnus aff. glutinosa subsp. barbata. h: Oriolo MSF 852. i: Oriolo MSF 850. j: Seed cone, Oriolo MSF 987. k–o: Corylus aff. avellana. k: Leaf, Oriolo MSF 841. l: Fruit, Oriolo MSF 1001. m: Fruit, Oriolo MSF 1022. n: Fruit, Oriolo MSF 1003. o: Fruit, Oriolo MSF 1022. Scale bars 50 mm (a–e, g–i), 10 mm (f, j–o).