Zelkova zelkovifolia (UNGER) BŮŽEK et KOTL., 1963

Denk, Thomas, Sami, Marco, Teodoridis, Vasilis & Martinetto, Edoardo, 2022, The Late Early Pleistocene Flora Of Oriolo, Faenza (Italy): Assembly Of The Modern Forest Biome, Fossil Imprint 78 (1), pp. 217-262 : 230

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https://doi.org/ 10.37520/fi.2022.009

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Zelkova zelkovifolia (UNGER) BŮŽEK et KOTL., 1963


Zelkova zelkovifolia (UNGER) BŮŽEK et KOTL., 1963

Text-figs 6m View Text-fig , 7a–d View Text-fig

1847 Ulmus zelkovifolia UNGER , p. 94 pro parte, pl. 24, figs 9–13, not fig. 8.

1963 Zelkova zelkovifolia (UNGER) BŮŽEK et KOTL. in Kotlaba, p. 59, pl. 3, figs 7, 8.

M a t e r i a l. Oriolo MSF 633, 639, 675, 681?, 682-1, 683, 685?, 811, 812, 813, 814, 815, 816, 817?, 818, 819-1, 820, 850; Tebano MSF n.n.; Santa Lucia delle Spianate MSF n.n.

D e s c r i p t i o n. Leaf, petiolate, lamina elliptical to ovate, lamina length 25–49 mm, width 16–25 mm, apex acute, base obtuse, sometimes slightly asymmetrical, margin serrate with coarse teeth (appearing crenate in case of blunt tooth apices), secondary venation craspedodromous, secondary veins commonly branching.

R e m a r k s. The leaves of Oriolo are similar to both the modern Z. carpinifolia (PALLAS) K.KOCH, 1849 , ranging from Turkey to northern Iran, and to the narrowly endemic Z. sicula DI PASQ., GARFI et QUÉZEL, 1992 , native to small areas in Sicily. For example, the leaf figured in Text-fig. 7d View Text-fig matches well leaves of Z. carpinifolia , while the leaf in Text-fig. 7b View Text-fig closely resembles leaves of Z. sicula ( Denk and Grimm 2005) . This last fossil is extremely similar to two fossil leaves from Riano Romano, figured by Follieri et al. (1986). Molecular data indicate that Z. sicula might be the result of a relatively recent hybridisation, with parents close to the ancestral species of Z. abelicea (LAM.) BOISS. (Crete) and the widespread Euxinian-Hyrcanian Z. carpinifolia ( Christe et al. 2014, Rucinska 2012). Roiron (1983) referred Zelkova leaves of the Plio-Pleistocene of Crespià to the modern species Z. crenata SPACH., 1841 (= Z. carpinifolia ). Because the present leaves show a mosaic of characteristics typical of two modern species, we consider them to represent the fossil-taxon Z. zelkovifolia .













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