Landouria madurensis, Nurinsiyah & Neiber & Hausdorf, 2019

Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, European Journal of Taxonomy 526, pp. 1-73 : 41-42

publication ID

https://doi.org/ 10.5852/ejt.2019.526

publication LSID

lsid:zoobank.org:pub:3FFC527D-E9CA-4D9F-BF59-076FA60171AC

DOI

https://doi.org/10.5281/zenodo.10447785

persistent identifier

https://treatment.plazi.org/id/024A8B78-91C2-49CA-8721-C0B8CCC46982

taxon LSID

lsid:zoobank.org:act:024A8B78-91C2-49CA-8721-C0B8CCC46982

treatment provided by

Plazi

scientific name

Landouria madurensis
status

sp. nov.

Landouria madurensis View in CoL sp. nov.

urn:lsid:zoobank.org:act:024A8B78-91C2-49CA-8721-C0B8CCC46982

Figs 8 View Figs 6–11 , 22 View Figs 15–24 , 36 View Figs 29–47 , 53 View Figs 52–55 , 57 View Figs 56–59 ; Tables 3–4 View Table 3 View Table 4

Helix squamulosa Martens, 1867: 266 , pl. 13, fig. 10 (“ Insel Madura (an der Nordostseite von Java), bei Sumanap”) (objective junior synonym of Helix squamulosa Mousson, 1856 ). Syn. nov.

Helix squamulosa – Pfeiffer 1871 in 1870–1876: pl. 117, figs 13–15; 1872 in 1870–1876: 41. Landouria squamulosa – van Benthem Jutting 1941: 311 .

Diagnosis

Landouria madurensis sp. nov. is characterized by a sharply keeled shell with longish scales all over the teleoconch, a wide umbilicus, a long penis, which is divided into a longer, broader distal and a shorter, narrower proximal part, and a long flagellum, which gradually tapers towards its proximal end.

Etymology

Landouria madurensis sp. nov. is named after its occurrence on Madura Island.

Material examined

Holotype

INDONESIA • holotype (det. anat.); East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest; 6°54′01″ S, 113°12′08″ E; 17 m a.s.l.; 14 May 2014; A.S. Nurinsiyah and T. Setiadi leg.; D = 11.4 mm, H = 6.5 mm; MZB 21432.

GoogleMaps

Paratypes

INDONESIA • 1 spec.; same data as for holotype; MZB 19209 • 4 spec.; same data as for holotype; MZB 19210 GoogleMaps 1 spec.; same data as for holotype; ZMH 133408 View Materials GoogleMaps 4 spec.; same data as for holotype; ZMH 133437 View Materials GoogleMaps 2 spec. (also syntypes of H. squamulosa ); Sumenep (see Martens 1867: 267); 7°01′ S, 113°52′ E; ZMB 7155 View Materials GoogleMaps .

GoogleMaps

Description

SHELL ( Figs 8 View Figs 6–11 , 57; Tables 3–4 View Table 3 View Table 4 ). Depressed conical, with 5.5–5.75 hardly convex whorls; protoconch almost smooth; teleoconch with irregular wrinkles; without distinct incised spiral lines; longish scales all over teleoconch; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture rounded rhombic; upper insertion of peristome descending; peristome expanded, reflexed and thickened; umbilicus slightly eccentric, very wide, comprising 38–40% of shell diameter, hardly obscured by columellar edge.

GENITALIA ( Figs 22 View Figs 15–24 , 36 View Figs 29–47 ; Table 4 View Table 4 ). Atrium short; penis long, divided into a longer, broader distal and a shorter, narrower proximal part, without penis coecum; membranaceous sheath around distal part of penis connected with proximal part of epiphallus by tissue; penial retractor runs from diaphragm to middle part of cylindrical epiphallus; flagellum long, broadest in its middle part, gradually tapering towards its proximal end, slightly crenated, without node; vas deferens narrow and long; vagina cylindrical, long; oviduct shorter, proximally hardly tapering; peduncle of bursa copulatrix subdivided into a broader, cylindrical distal part and a narrower, longer proximal part. Right ommatophoral retractor runs between penis and vagina.

Remarks

Landouria madurensis sp. nov. differs from the neighbouring Landouria intumescens in the presence of longish scales all over the teleoconch ( Fig. 8 View Figs 6–11 ), a wider umbilicus (U/D 0.38–0.40 vs 0.22–0.34 in L. intumescens ) and a relatively long vagina (P:V 2.1 vs 2.4–3.9 in L. intumescens ).

A second, more conical species of Landouria without longish scales and with a narrower umbilicus, of which only juveniles were found, lives sympatrically with Landouria madurensis sp. nov. in Nepa Monkey lowland rainforest on Madura Island.

Distribution

Landouria madurensis sp. nov. is endemic to Madura Island (Fig. 53).

ZMH

USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)

MZB

Museum Zoologicum Bogoriense

Kingdom

Animalia

Phylum

Mollusca

Class

Gastropoda

Order

Stylommatophora

Family

Camaenidae

Genus

Landouria

Loc

Landouria madurensis

Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard 2019
2019
Loc

Helix squamulosa –

van Benthem Jutting W. S. S. 1941: 311
1941
Loc

Helix squamulosa

Martens E. von 1867: 266
1867
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