Landouria smimensis ( Mousson, 1848 )

Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, European Journal of Taxonomy 526, pp. 1-73 : 57-58

publication ID

https://doi.org/ 10.5852/ejt.2019.526

publication LSID

lsid:zoobank.org:pub:3FFC527D-E9CA-4D9F-BF59-076FA60171AC

DOI

https://doi.org/10.5281/zenodo.11238091

persistent identifier

https://treatment.plazi.org/id/03B1DB4C-8B15-4E20-4C1A-FE2B3DF9FBCE

treatment provided by

Plazi

scientific name

Landouria smimensis ( Mousson, 1848 )
status

 

Landouria smimensis ( Mousson, 1848) View in CoL

Figs 11 View Figs 6–11 , 43 View Figs 29–47 , 52 View Figs 52–55 , 67 View Figs 63–71 , 78–80 View Figs 76–80 ; Tables 3–4 View Table 3 View Table 4

Helix smimensis Mousson, 1848: 266 View in CoL (“ Java ”; more exactly ( Mousson 1849): “Smim, 4000′”).

Helix smironensis Mousson, 1850: 177 View in CoL (unjustified emendation).

Helix smiruensis Martens, 1867: 268 View in CoL (unjustified emendation).

Plectotropis tenggerica Schepman, 1912: 233 , pl. 10, figs 9–11 (“Nongkodjadjar”).

Helix smimensis – Mousson 1849: 21 View in CoL , pl. 2, fig. 10.

Plectotropis View in CoL sp. – Schepman 1915: 19.

Landouria smironensis – van Benthem Jutting 1950: 467 View in CoL (in part).

Diagnosis

Landouria smimensis is characterized by a small, depressed conical shell with hairs and a usually slightly angular body whorl, a long penis, which gradually tapers towards its proximal end, and a slightly crenated, gradually tapering flagellum.

Material examined

Type material

INDONESIA – East Java • holotype of P. tenggerica ; Nongkojajar ; 7°54′ S, 112°49′ E; ZMA 174453 View Materials GoogleMaps 2 syntypes of H. smimensis ; Gunung Semeru ; 8°06′ S, 112°55′ E; ca 1200 m a.s.l., ZMZ 508598 GoogleMaps .

Other material

INDONESIA • 23 spec. (vouchers for Martens 1867); Java; ZMB 4062 View Materials 1 spec. (voucher for van Benthem Jutting 1950); Java; ZMB 67043 . – East Java • 3 spec. (vouchers for van Benthem Jutting 1950); Nongkojajar , near waterfall ; 7°54′ S, 112°49′ E; ZMA 407685 View Materials GoogleMaps many spec. (vouchers for van Benthem Jutting 1950); same data as for preceding; ZMA 407681 View Materials GoogleMaps 7 spec.; Nongkojajar, near Baron ; 7°54′ S, 112°49′ E; ZMA 407686 View Materials GoogleMaps 2 spec. (vouchers for van Benthem Jutting 1950) (det. anat.); Tosari ; 7°53′ S, 112°54′ E; ca 1800 m a.s.l.; ZMA 390881 View Materials GoogleMaps .

Description

SHELL ( Figs 11 View Figs 6–11 , 78–80 View Figs 76–80 ; Tables 3–4 View Table 3 View Table 4 ). Depressed conical, with 4.5–5.5 convex whorls; protoconch almost smooth; teleoconch irregularly striated, with indistinct incised spiral lines; with hair-like processes; not tuberculate around umbilicus; brownish-corneous; body whorl slightly angular or rounded; aperture oval; upper insertion of peristome hardly descending; peristome sharp; umbilicus slightly eccentric, narrow, comprising 18–32% of shell diameter, hardly obscured by columellar edge.

GENITALIA ( Figs 43 View Figs 29–47 , 67 View Figs 63–71 ; Table 4 View Table 4 ). Atrium short; penis long, gradually tapering towards its proximal end; without penis coecum; surrounded by a membranaceous sheath connected to proximal epiphallus by tissue; penial retractor runs from diaphragm to middle part of cylindrical epiphallus; flagellum moderately long, narrow, broadest in its middle part, without node, gradually tapering towards its proximal end, slightly crenated; vas deferens narrow and long; vagina broad and short; oviduct short, proximally slightly tapering; peduncle of bursa copulatrix subdivided into a broad abruptly tapering distal part and a narrower and longer proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.

Remarks

The original spelling of the species name was Helix smimensis Mousson, 1848 . This was not an inadvertent error, such as a lapsus calami or printer’s error, because this spelling was also used later by Mousson (1849). Thus, H. smimensis is the correct original spelling according to ICZN, Article 32. The name was intentionally changed to H. smironensis by Mousson (1850) and to H. smiruensis by Martens (1867). Both names are unjustified emendations, which are available with these authors and dates, and which are junior objective synonyms of H. smimensis (ICZN, Art. 33). According to ICZN, Article 33.2.3.1, an unjustified emendation may be deemed to be a justified emendation, if it is in prevailing usage. Helix smiruensis was used by Tryon (1887), Leschke (1914) and Rensch (1932), L. smironensis by van Benthem Jutting (1950) and Vermeulen & Whitten (1998), the incorrect subsequent spelling Landouria cf. smiroensis by Boonngam et al. (2008) and the correct original spelling L. smimensis by Köhler et al. (in press). Thus, there is no prevailing usage of any of the emendations and the correct original spelling should be used.

The voucher specimen for L. tenggerica of Rensch (1934) from Gunung Lawu, 2400 m a.s.l. (ZMB 76086) is tuberculate around the umbilicus and is a juvenile of a different species.

Distribution

Landouria smimensis View in CoL is known from the Tengger Mountains in East Java (Fig. 52). It has also been recorded from other mountains in Central and East Java ( Rensch 1934), Lombok ( Rensch 1932), Bali ( Vermeulen & Whitten 1998) and Thailand ( Boonngam et al. 2008), but these populations probably represent separate species.

ZMB

Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet

ZMB

Museum für Naturkunde Berlin (Zoological Collections)

Kingdom

Animalia

Phylum

Mollusca

Class

Gastropoda

Order

Stylommatophora

Family

Camaenidae

Genus

Landouria

Loc

Landouria smimensis ( Mousson, 1848 )

Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard 2019
2019
Loc

Landouria smironensis – van Benthem Jutting 1950: 467

van Benthem Jutting W. S. S. 1950: 467
1950
Loc

Plectotropis

Schepman M. M. 1915: 19
1915
Loc

Plectotropis tenggerica

Schepman M. M. 1912: 233
1912
Loc

Helix smiruensis

Martens E. von 1867: 268
1867
Loc

Helix smironensis

Mousson A. 1850: 177
1850
Loc

Helix smimensis –

Mousson A. 1849: 21
1849
Loc

Helix smimensis

Mousson A. 1848: 266
1848
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